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Coleoptera sp.



Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordines: Coleopterida – Hymenopterida – Neuropterida – Panorpida
incertae sedis: †Cavalarva – †Metabolarva

Overview of ordines


Coleoptera – Diptera – Hymenoptera – Lepidoptera – "Mecoptera" – Megaloptera – Neuroptera – Raphidioptera – Siphonaptera – Strepsiptera – Trichoptera – †Permotrichoptera – †Skleroptera
[list of ordines after Grimaldi & Engel (2005: 147, table 4.1)]



Holometabola Burmeister, 1835


  • Metabola Burmeister, 1832
  • Holometabola Burmeister, 1835
  • Endopterygota Sharp, 1898



Primary references

  • Sharp, D. 1898. Some points in the classification of Insecta Hexapoda. Pp. 246–249. In: Sedgwick, A. (ed.) Proceedings of the fourth International Congress of Zoology, Cambridge, 22-27 August, 1898. London: C.J. Clay and Sons. BHL.

Additional references

  • Aristov, D.S. & Rasnitsyn, A.P. 2015. New insects from the Kungurian of Tshekarda fossil site in Permian Territory of Russia. Russian Entomological Journal 24(1): 17–35. DOI: 10.15298/rusentj.24.1.03 Open access. Reference page
  • Aspöck, U. 2002: Phylogeny of the Neuropterida (Insecta: Holometabola). Zoologica Scripta, 31: 51–56.
  • Beutel, R.G. and H. Pohl. 2006: Endopterygote systematics – where do we stand and what is the goal (Hexapoda, Arthropoda)? Systematic Entomology, 31(2): 202–219.
  • Büning, J. 2005: The telotrophic ovary known from Neuropterida exists also in the myxophagan beetle Hydroscapha natans. Development genes and evolution, 215: 597–607.
  • Büning, J. 2006: Ovariole structure supports sistergroup relationship of Neuropterida and Coleoptera. Arthropod systematics & phylogeny, 64: 115–126. [1]
  • Castro, L.R.; Dowton, M. 2005: The position of the Hymenoptera within the Holometabola as inferred from the mitochondrial genome of Perga condei (Hymenoptera: Symphyta: Pergidae). Molecular Phylogenetics and Evolution, 34(3): 469–479.
  • Grimaldi, D. & Engel, M.S. 2005. Evolution of the Insects. Cambridge University Press, New York, USA. ISBN 978-0-521-82149-0. limited preview on Google Books. Reference page
  • Haring, E.; Aspöck, U. 2004: Phylogeny of the Neuropterida: a first molecular approach. Systematic entomology, 29: 415–430.
  • Krenn, H.W. 2007: Evidence from mouthpart structure on interordinal relationships in Endopterygota? Arthropod systematics & phylogeny, 65(1): 7–14. PDF
  • Nel, A., Roques, P., Nel, P., Prokin, A.A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D.-Y., Engel, M. & Kirejtshuk, A.G. 2013. The earliest-known holometabolous insects. Nature 503: 257–261. [14 p. Supplementary material.] DOI: 10.1038/nature12629 Paywall. Reference page
  • Wootton, R.J. 2002: Design, function and evolution in the wings of holometabolous insects. Zoologica Scripta, 31: 31–40.
  • Whiting, M.F. 2002: Phylogeny of the holometabolous insect orders: molecular evidence. Zoologica Scripta, 31: 3–15.

Vernacular names

English: Complete Metamorphosis Insects
magyar: Fejlett szárnyas rovarok
日本語: 内翅類
Türkçe: Tümbaşkalaşmalı böcekler
Tiếng Việt: Côn trùng trưởng thành
中文: 内翅總目
Wikimedia Commons For more multimedia, look at Holometabola on Wikimedia Commons.