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Taxonavigation: Scarabaeoidea 

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Phanaeini
Genera (12): Bolbites – Coprophanaeus – Dendropaemon – Diabroctis – Gromphas – Homalotarsus – Megatharsis – Oruscatus – Oxysternon – Phanaeus – Sulcophanaeus – Tetramereia



Phanaeini MacLeay, 1819


  • Macleay, W.S. 1819. Horae entomologicae: or essays on the annulose animals. Containing geneal observations on the geography, manners, and natural affinities of the Insects which compose the genus Scarabaeus of Linnaeus; to which are added a few incidental remarks on the genera Lucanus and Hister of the same author. With an appendix and plates. Vol. 1. Part 1. London: S. Bagster, xxx + [2] + 160 pp., 3 pls. BHL Reference page
  • Gillett, C.P.D.T.; Gillett, M.P.T.; Gillett, J.E.D.T.; Vaz-de-Mello, F.Z. 2010: Diversity and distribution of the scarab beetle tribe Phanaeini in the northern states of the Brazilian Northeast (Coleoptera: Scarabaeidae: Scarabaeinae). Insecta mundi, (0118 ): 1–19. PDF
  • Lizardo, V., Escobar, F. & Rojas-Soto, O. 2017. Diversity and distribution of Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Mexico. Zootaxa 4358(2): 271–294. DOI: 10.11646/zootaxa.4358.2.3. Reference page
  • Puker, A.; Correa, C.M.A.; Korasaki, V. 2014: Deltochilini and Phanaeini dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in introduced and native ecosystems of Brazil. Journal of natural history, DOI: 10.1080/00222933.2014.908969 Reference page