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Taxonavigation: Scarabaeoidea 

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Scarabaeini
Genus: Pachysoma
Species (15): P. aesculapius – P. bennigseni – P. denticolle – P. endroedyi – P. fitzsimonsi – P. gariepinum – P. glentoni – P. hessi – P. hippocrates – P. marginatum – P. rodriguesi – P. rotundigenum – P. schinzi – P. striatum – P. valeflorae


Pachysoma Macleay, 1821


[list of synonyms after Harrison, Scholtz & Chown (2003: 311)]


Additional references[edit]

  • Forgie, S.A.; Kryger, U.; Bloomer, P.; Scholtz, C.H. 2006: Evolutionary relationships among the Scarabaeini (Coleoptera: Scarabaeidae) based on combined molecular and morphological data. Molecular phylogenetics and evolution, 40: 662–678. ISSN: 1055-7903 DOI: 10.1016/j.ympev.2006.04.018
  • Harrison, J. du G.; Philips, T.K. 2003: Phylogeny of Scarabaeus (Pachysoma MacLeay) stat. nov., and related flightless Scarabaeini (Scarabaeidae: Scarabaeinae). Annals of the Transvaal Museum 40: 47–71. full article (PDF) Reference page
  • Harrison, J.du G.; Scholtz, C.H.; Chown, S.L. 2003: A revision of the endemic south-western African dung beetle subgenus Scarabaeus (Pachysoma) MacLeay, including notes on other flightless Scarabaeini (Scarabaeidae: Scarabaeinae). Journal of natural history, 37: 305–355. DOI: 10.1080/713834683
  • Holm, E.; Scholtz, C.H. 1979: A revision of the genus Pachysoma M'Leay with an evaluation of the subtribe Pachysomina Ferreira and its genera (Coleoptera: Scarabaeidae). Journal of the Entomological Society of Southern Africa, 42 (2): 225–244.
  • Scholtz, C.H.; Harrison, J. du G.; Grebennikov, V.V. : Dung beetle (Scarabaeus (Pachysoma)) biology and immature stages: reversal to ancestral states under desert conditions (Coleoptera: Scarabaeidae)? Biological journal of the Linnean Society, 83 (4): 453–460. DOI: 10.1111/j.1095-8312.2004.00389.x
  • Sole, C.L.; Scholtz, C.H.; Bastos, A.D.S. 2005: Phylogeography of the Namib Desert dung beetles Scarabaeus (Pachysoma) MacLeay (Coleoptera: Scarabaeidae). Journal of biogeography, 32: 75–84. DOI: 10.1111/j.1365-2699.2004.01124.x


  • Schoolmeesters P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page.  [accessed on April 4, 2017]