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Taxonavigation: Scarabaeoidea 

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Orphninae
Tribus (2): Aegidiini - Orphnini

Overview of genera (15)[edit]

Aegidiellus – Aegidinus – Aegidium – Chaetonyx – Craniorphnus – Goniorphnus – Hybaloides – Hybalus – Madecorphnus – Orphnus – Paraegidium – Pseudorphnus – Renorphnus – Stenosternus – Triodontus
[source: Frolov (2012)]


  • Colby, J. 2009: Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi, 0076: 1–41. PDF
  • Frolov, A.V. 2008: A new apterous species of the genus Orphnus MacLeay (Coleoptera: Scarabaeidae: Orphninae) from South Africa. Zootaxa, 1855: 65–68. Abstract & excerpt
  • Frolov, A.V. 2009: New scarab-beetle species (Coleoptera, Scarabaeidae, Aphodiinae, Orphninae) from central Asia and southern Africa. Entomological review, 89: 685–688. DOI: 10.1134/S0013873809060062
  • Frolov, A.V. 2010: Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera: Scarabaeidae: Orphninae). Journal of natural history, 44: 1095–1111. DOI: 10.1080/00222931003632765
  • Frolov, A.V. 2010: [A new species of the genus Triodontus (Coleoptera, Scarabaeidae, Orphninae) from Southern Madagascar.] Entomologicheskoe obozrenie, 89 (4): 835–839. [not seen, in Russian, English translation in Entomological review, 90 (8): 1031–1034. (2010) DOI: 10.1134/S0013873810080087 ]
  • Frolov, A.V. 2012: Diagnosis, classification, and phylogenetic relationships of the orphnine scarab beetles (Coleoptera, Scarabaeidae: Orphninae). Entomologicheskoe obozrenie, 91(2): 332–351. [in Russian, English translation in Entomological review, 92(7): 782–797. (2012) DOI: 10.1134/S0013873812070056 ] Reference page
  • Frolov, A.V. 2013: Little known Orphninae (Coleoptera, Scarabaeidae) collected by the Madagascan expeditions of the California Academy of Sciences. Biodiversity data journal, 1: e1016. DOI: 10.3897/BDJ.1.e1016 Reference page
  • Frolov, A.V.; Montreuil O. 2009: A new genus of Orphninae (Coleoptera: Scarabaeidae) from Madagascar. Zoosystematica Rossica, 18: 65–69.