Iolaus frater

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Epamera frater, ♂, from original description.


Taxonavigation: Papilionoidea 

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Lycaenidae
Subfamilia: Theclinae
Tribus: Iolaini
Genus: Iolaus
Subgenus: Iolaus (Epamera)
Species: Iolaus frater
I. f. frater – I. f. kamerunica – I. f. kumboae


Iolaus frater (Joicey & Talbot, 1921).

Type locality: Democratic Republic of Congo: “Between Lindi and Lubila Rivers, N. of Batama”.

Syntypes: 3♂♂ BMNH, iv.1920 (T.A. Barns).


  • Epamera frater Joicey & Talbot, 1921: 92, pl.15, figs 68,69.
  • Epamera pater [sic] Joicey & Talbot; Riley, 1928: 384, [mis-spelling].
  • Iolaus (Epamera) frater (Joicey & Talbot); Stempffer, 1967: 146; Williams, 2007: Iolaina 41.