Carabus variolosus nodulosus

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The status, affinity, scope or nomenclature of this taxon is disputed.

Please see discussion on the talk page.


Taxonavigation: Adephaga 

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Carabidae
Subfamilia: Carabinae
Tribus: Carabini
Genus: Carabus
Division: Carabus div. Multistriati
Subdivision: Carabus subdiv. Procrustimorphi
Subgenus: Carabus (Hygrocarabus)
Species: Carabus variolosus
Subspecies: Carabus variolosus nodulosus


  • Carabus variolosus nodulosus Creutzer, 1799



  • Matern, A.; Drees, C.; Kleinwächter, M.; Assmann, T. 2007: Habitat modelling for the conservation of the rare ground beetle species Carabus variolosus (Coleoptera, Carabidae) in the riparian zones of headwaters. Biological conservation, 136(4): 618–627. DOI: 10.1016/j.biocon.2007.01.006
  • Matern, A.; Drees, C.; Meyer, H.; Assmann, T. 2008: Population ecology of the rare carabid beetle Carabus variolosus (Coleoptera: Carabidae) in north-west Germany. Journal of insect conservation, 12(6): 591–601. DOI: 10.1007/s10841-007-9096-3