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Taxonavigation: Caryophyllales 
Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Ordo: Caryophyllales

Familia: Caryophyllaceae
Tribus: Sperguleae
Genus: Rhodalsine – Sanctambrosia – Spergula – Spergularia


Sperguleae Dumort., Anal. Fam. Pl.: 49. (1829)

Note: Kool (2012) confirmed that Spergula and Spergularia, as circumscribed on their taxon pages, are separate monophyletic genera.


  • Homotypic
    • Spergulinae Dumort., Fl. Belg. 109. (1827)
    • Sperguloideae Darby, Man. Bot. vii. (1841)
    • Spergularieae Hartm., Handb. Skand. Fl., ed. 5: 137. (1849)


  • Dumortier, B.C.J. 1829. Analyse des Familles de Plantes: avec l'indication des principaux genres qui s'y rattachent. Tournay 49.
  • Dillenberger, M.S. & Kadereit, J.W. 2014. Maximum polyphyly: Multiple origins and delimitation with plesiomorphic characters require a new circumscription of Minuartia (Caryophyllaceae). Taxon 63(1): 64–88. doi: 10.12705/631.5 Reference page. 
  • Greenberg, A.K. & Donoghue, M.J. 2011. Molecular systematics and character evolution in Caryophyllaceae. Taxon 60(6): 1637–1652. Full text PDF reference page .
  • Harbaugh, D.T., Nepokroeff, M., Rabeler, R.K., McNeill, J., Zimmer, E.A. & Wagner, W.L. 2010. A new lineage based tribal classification of the family Caryophyllaceae. International Journal of Plant Sciences 171(2): 185–198. doi: 10.1086/648993 Full text from Research Gate reference page 
  • Hernández-Ledesma, P., Berendsohn, W. G., Borsch, T., Mering, S. von, Akhani, H., Arias, S., Castañeda-Noa, I., Eggli, U., Eriksson, R., Flores-Olvera, H., Fuentes-Bazán, S., Kadereit, G., Klak, C., Korotkova, N., Nyffeler R., Ocampo G., Ochoterena, H., Oxelman, B., Rabeler, R. K., Sanchez, A., Schlumpberger, B. O. & Uotila, P. 2015. A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales. Willdenowia 45(3): 281–383. doi: 10.3372/wi.45.45301 Full text PDF Reference page. 
  • Kool, A. 2012. Desert plants and deserted islands: systematics and ethnobotany in Caryophyllaceae. Doctoral dissertation Full text available, Acta Universitatis Upsaliensis. Reference page. 
  • Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website. Version 12, July 2012 [and more or less continuously updated since]. (web site)