Template:Wasshausen, 1995
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- Wasshausen, D.C. 1995. Acanthaceae. Pp. 335–374 in Steyermark, J.A. (†), Berry, P.E. & Holst, B.K. (eds.), Flora of the Venezuelan Guayana, Vol. 2, Pteridophytes, Spermatophytes, Acanthaceae–Araceae. Missouri Botanical Garden Press, St. Louis. ISBN 0-915279-74-6.