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Ochi, Kawahara & Masumoto, 2019
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From Wikispecies
Ochi, T.
,
Kawahara, M
&
Masumoto, K.
2019. Two new subspecies of Onthophagini species from the Ryukyu Islands, Southwestern Islands of Japan (Coleoptera, Scarabaeidae).
Kogane
22: 38-42.
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Ochi, Kawahara & Masumoto, 2019
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