Template:Howden, 1978
- Howden, H.F. 1978. Descriptions of some West Indian Scarabaeidae primarily in the Natural History Museum, Basel. Entomologica Basiliensia 3: 377-394. [https://www.e-periodica.ch/cntmng?pid=enc-001%3A1978%3A3%3A%3A654 Full articel (PDF).