Template:González-Bergonzoni et al., 2009
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- González-Bergonzoni, I., Loureiro, M. & Oviedo, S. 2009. A new species of Gymnogeophagus from the río Negro and río Tacuarí basins, Uruguay (Teleostei: Perciformes). Neotropical Ichthyology 7(1): 19–24.