Template:Gebien, 1922a
- Gebien, H. 1922. Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner. No. V. Coleoptera, Heteromera: Tenebrionidae. Transactions of the Linnean Society of London 2 Series, Zoology. 18: 261–324, pl. 23.