Template:Erwin, 2004
- Erwin, T.L. 2004. The beetle family Carabidae of Costa Rica and Panamá: descriptions of four new genera and six new species with notes on their way of life (Insecta: Coleoptera). Zootaxa 537: 1–18. DOI: 10.11646/zootaxa.537.1.1, PDF