Template:Emmel,Emmel,Mattoon, 1998 36
- Emmel, J.F., Emmel, T. & Mattoon, S., 1998. A New Subspecies of Speyeria zerene (Lepidoptera: Nymphalidae) from the San Francisco Bay Area of California. In Systematics of Western North American Butterflies. 36: 455-460.