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This is the basal grade of Polyphaga, according to Hunt et al. (2007). Their proposed phylogeny is:

((Scirtidae , Decliniiidae ), (Derodontidae , ((Eucinetidae , Clambidae ), remaining Polyphaga)))

See discussion in Maddison et al. (2008: 17).


  • Hunt, T.; Bergsten, J.; Levkanicova, Z.; Papadopoulou, A.; St. John, O.; Wild, R.; Hammond, P.M.; Ahrens, D.; Balke, M.; Caterino, M.S.; Gomez-Zurita, J.; Ribera, I.; Barraclough, T.G.; Bocakova, M.; Bocak, L.; Vogler, A.P. 2007: A comprehensive phylogeny of beetles reveals the evolutionary origins of a superradiation. Science, 318: 1913-1916. [1]
  • Maddison, D.R.; Moore, W.; Baker, M.D.; Ellis, T.M.; Ober, K.A.; Cannone, J.J.; Gutell, R.R. 2008: Monophyly of terrestrial adephagan beetles as indicated by three nuclear genes (Coleoptera: Carabidae and Trachypachidae). Zoologica scripta, doi: 10.1111/j.1463-6409.2008.00359.x

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