Robert Alexander Pyron

Robert Alexander Pyron, U.S. herpetologist.

• Department of Biological Sciences, The George Washington University, Washington D.C., United States of America.

Taxon names authored

(List may be incomplete)

• 63 taxon names authored by Robert Alexander Pyron

Publications

(List may be incomplete)

2008

• Burbrink, F.T., Fontanella, F., Pyron, R.A., Guiher, T.J. & Jimenez, C. 2008. Phylogeography across a continent: The evolutionary and demographic history of the North American racer (Serpentes: Colubridae: Coluber constrictor). Molecular Phylogenetics and Evolution 47(1): 274–288. DOI: 10.1016/j.ympev.2007.10.020 Reference page. 

2011

• Pyron, R.A. & Wiens, J.J. 2011. A large-scale phylogeny of Amphibia including over 2,800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61(2): 543–583. DOI: 10.1016/j.ympev.2011.06.012 Reference page. 
• Pyron, R.A., Burbrink, F.T., Colli, G.R., Nieto-Montes de Oca, A., Vitt, L.J., Kuczynski, C.A. & Wiens, J.J. 2011. The phylogeny of advanced snakes (Colubroidea), with discovery of a new subfamily and comparison of support methods for likelihood trees. Molecular Phylogenetics and Evolution 58(2): 329–342. DOI: 10.1016/j.ympev.2010.11.006 Reference page. 

2012

• Burbrink, F.T., Ruane, S. & Pyron, R.A. 2012. When are adaptive radiations replicated in areas? Ecological opportunity and unexceptional diversification in West Indian dipsadine snakes (Colubridae: Alsophiini). Journal of Biogeography 39(3): 465–475. DOI: 10.1111/j.1365-2699.2011.02621.x Reference page. 

2013

• Abraham, R.K., Pyron, R.A., Ansil, B.R., Zachariah, A. & Zachariah, A. 2013. Two novel genera and one new species of treefrog (Anura: Rhacophoridae) highlight cryptic diversity in the Western Ghats of India. Zootaxa 3640(2): 177–199. DOI: 10.11646/zootaxa.3640.2.3 Reference page. 
• Guo, P., Zhang, L., Liu, Q., Li, C., Pyron, R.A., Jiang, K. & Burbrink, F.T. 2013. Lycodon and Dinodon: one genus or two? Evidence from molecular phylogenetics and morphological comparisons. Molecular Phylogenetics and Evolution 68(1): 144–149. DOI: 10.1016/j.ympev.2013.03.008 Reference page. 
• Pyron, R.A., Burbrink, F.T. & Wiens, J.J. 2013. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology 13(93): 1–53. DOI: 10.1186/1471-2148-13-93 . ResearchGate . Reference page. 

2014

• Pyron, R.A. & Wallach, V. 2014. Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence. Zootaxa 3829(1): 1–81. DOI: 10.11646/zootaxa.3829.1.1 Reference page. 
• Pyron, R.A., Reynolds , R.G. & Burbrink, F.T. 2014. A taxonomic revision of boas (Serpentes: Boidae). Zootaxa 3846(2): 249–260. DOI: 10.11646/zootaxa.3846.2.5 Reference page. 
• Guo, G., Zhu, F., Liu, Q., Zhang, L., Li, J.X., Huang, Y.Y. & Pyron, R.A. 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873(4): 425–440. DOI: 10.11646/zootaxa.3873.4.5 Reference page. 
• Ruane, S., Bryson, Jr., R.W., Pyron, R.A. & Burbrink, F.T. 2014. Coalescent Species Delimitation in Milksnakes (genus Lampropeltis) and Impacts on Phylogenetic Comparative Analyses. Systematic Biology 63(2): 231–250. DOI: 10.1093/sysbio/syt099 Reference page. 

2015

• Guayasamin, J.M., Rivera-Correa, M., Arteaga, A., Culebras, J., Bustamante, L., Pyron, R.A., Peñafiel, N., Morochz, C. & Hutter, C.R. 2015. Molecular phylogeny of stream treefrogs (Hylidae: Hyloscirtus bogotensis Group), with a new species from the Andes of Ecuador. Neotropical Biodiversity 1(1): 2–21. DOI: 10.1080/23766808.2015.1074407 Reference page. 
• Pyron, R.A., Guayasimin, J.M., Peñafiel, N., Bustamente, L. & Arteaga, A. 2015. Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador. ZooKeys 541: 109–147. DOI: 10.3897/zookeys.541.6058 Reference page. 

2016

• Arteaga, A., Pyron, R.A., Peñafiel, N., Romero-Barreto, P., Culebras, J., Bustamante, L., Yánez-Muñoz, M.H. & Guayasamin, J.M. 2016. Comparative phylogeography reveals cryptic diversity and repeated patterns of cladogenesis for amphibians and reptiles in northwestern Ecuador. PLoS One 11(4: e0151746): 1–40. DOI: 10.1371/journal.pone.0151746 Reference page. 
• Pyron, R.A., Arteaga, A., Echevarría, L.Y. & Torres-Carvajal, O. 2016. A revision and key for the tribe Diaphorolepidini (Serpentes: Dipsadidae) and checklist for the genus Synophis. Zootaxa 4171(2): 293–320. DOI: 10.11646/zootaxa.4171.2.4. Reference page. 
• Pyron, R.A., Ganesh, S.R., Sayyed, A., Sharma, V., Wallach, V. & Somaweera, R. 2016. A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae). Zoosystema 38(4): 453–506. DOI: 10.5252/z2016n4a2 Reference page. 
• Pyron, R.A., Hsieh, F.W., Lemmon, A.R., Lemmon, E.M. & Hendry, C.R. 2016. Integrating phylogenomic and morphological data to assess candidate species-delimitation models in brown and red-bellied snakes (Storeria). Zoological Journal of the Linnean Society 177(4): 937–949. DOI: 10.1111/zoj.12392 Reference page. 
• Murphy, J.C., Jowers, M.J., Lehtinen, R.M., Charles, S.P., Colli, G.R., Péres Jr., A.K., Hendry, C.R. & Pyron, R.A. 2016. Cryptic, Sympatric Diversity in Tegu Lizards of the Tupinambis teguixin Group (Squamata, Sauria, Teiidae) and the Description of Three New Species. PLoS ONE 11(8): e0158542. DOI: 10.1371/journal.pone.0158542 Reference page. 
• Sayyed, A., Pyron, R.A. & Dahanukar, N. 2016. Cnemaspis flaviventralis, a new species of gecko (Squamata: Gekkonidae) from the Western Ghats of Maharashtra, India. Journal of Threatened Taxa 8(14): 9619–9629. DOI: 10.11609/jott.2599.8.14.9619-9629 Reference page. 
• Tucker, D.B., Colli, G.R., Giugliano, L.G., Hedges, S.B., Hendry, C.R., Lemmon, E.M., Lemmon, A.R., Sites Jr., J.W. & Pyron, R.A. 2016. Methodological congruence in phylogenomic analyses with morphological support for teiid lizards (Sauria: Teiidae). Molecular Phylogenetics and Evolution 103: 75–84. DOI: 10.1016/j.ympev.2016.07.002 . Reference page. 

2017

• Pyron, R.A. 2017. Novel Approaches for Phylogenetic Inference from Morphological Data and Total-Evidence Dating in Squamate Reptiles (Lizards, Snakes, and Amphisbaenians). Systematic Biology 66(1): 38–56. DOI: 10.1093/sysbio/syw068 Reference page. 
• Chen, X., Lemmon, A.R., Lemmon, E.M., Pyron, R.A. & Burbrink, F.T. 2017. Using phylogenomics to understand the link between biogeographic origins and regional diversification in ratsnakes. Molecular Phylogenetics and Evolution 111: 206–218. DOI: 10.1016/j.ympev.2017.03.017 Reference page. 
• Guayasamin, J.M., Hutter, C.R., Tapia, E.E., Culebras, J., Peñafiel, N., Pyron, R.A., Morochz, C., Funk, W.C. & Arteaga, A. 2017. Diversification of the rainfrog Pristimantis ornatissimus in the lowlands and Andean foothills of Ecuador. PLoS ONE 12(3): e0172615. DOI: 10.1371/journal.pone.0172615 Reference page. 
• Wickramasinghe, L.J.M., Vidanapathirana, D.R., Kandambi, H.K.D., Pyron, R.A. & Wickramasinghe, N. 2017. A New Species of Aspidura Wagler, 1830 (Squamata: Colubridae: Natricinae) from Sri Pada Sanctuary (Peak Wilderness), Sri Lanka. Zootaxa 4347(2): 275–292. DOI: 10.11646/zootaxa.4347.2.4. Reference page. 

2018

• Sayyed, A., Pyron, R.A. & Dileepkumar, R. 2018. Four new species of the genus Cnemaspis Strauch, (Sauria: Gekkonidae) from the northern Western Ghats, India. Amphibian & Reptile Conservation 12(2) [General Section]: 1–29 (e157). Reference page. 
• Arteaga, A., Salazar-Valenzuela, D., Mebert, K., Peñafiel, N., Aguiar, G., Sánchez-Nivicela, J.C., Pyron, R.A., Colston, T.J., Cisneros-Heredia, D.F., Yánez-Muñoz, M.H., Venegas, P.J., Guayasamin, J.M. & Torres-Carvajal, O. 2018. Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79–147. DOI: 10.3897/zookeys.766.24523 Reference page. 

2019

• Pyron, R.A. & Somaweera, R. 2019. Further notes on the Sri Lankan uropeltid snakes Rhinophis saffragamus (Kelaart, 1853) and Uropeltis ruhunae Deraniyagala, 1954. Zootaxa 4560(3): 592–600. DOI: 10.11646/zootaxa.4560.3.13 Reference page. 
• Vijayakumar, S.P., Pyron, R.A., Dinesh, K.P., Torsekar, V.R., Srikanthan, A.N., Swamy, P., Stanley, E.L., Blackburn, D.C. & Shanker, K. 2019. A new ancient lineage of frog (Anura: Nyctibatrachidae: Astrobatrachinae subfam. nov.) endemic to the Western Ghats of Peninsular India. PeerJ 7: e6457. DOI: 10.7717/peerj.6457 Reference page. 

2020

• Burbrink, F.T., Grazziotin, F.G., Pyron, R.A., Cundall, D., Donnellan, S., Irish, F., Keogh, J.S., Kraus, F., Murphy, R.W., Noonan, B., Raxworthy, C.J., Ruane, S., Lemmon, A.R., Lemmon, E.M. & Zaher, H. 2020. Interrogating Genomic-Scale Data for Squamata (Lizards, Snakes, and Amphisbaenians) Shows no Support for Key Traditional Morphological Relationships. Systematic Biology 69(3): 502–520. DOI: 10.1093/sysbio/syz062 Reference page. 
• Colston, T.J., Noonan, B.P., Smith, M.L. & Pyron, R.A. 2020. Availability of Recently Described Ptychadena (Anura: Ptychadenidae) Nomina from Ethiopia. Zootaxa 4786(2): 289–294. DOI: 10.11646/zootaxa.4786.2.11 Reference page. 
• Colston, T.J., Pyron, R.A. & Bauer, A.M. 2020. A new species of African Snake-Eyed Skink (Scincidae: Panaspis) from Ethiopia. Zootaxa 4779(2): 190–200. DOI: 10.11646/zootaxa.4779.2.2 Reference page. 
• Pyron, R.A. & Beamer, D.A. 2020. The herpetological legacy of Jacob Green and the nomenclature of some North American lizards and salamanders. Zootaxa 4838(2): 221–247. DOI: 10.11646/zootaxa.4838.2.4 Reference page. 
• Pyron, R.A., O’Connell, K.A., Lemmon, E.M., Lemmon, A.R. & Beamer, D.A. 2020. Phylogenomic data reveal reticulation and incongruence among mitochondrial candidate species in Dusky Salamanders (Desmognathus). Molecular Phylogenetics and Evolution 146: 106751. DOI: 10.1016/j.ympev.2020.106751 Reference page. 
• Rajabizadeh, M., Pyron, R.A., Nazarov, R., Poyarkov, N.A., Adriaens, D. & Herrel, A. 2020. Additions to the phylogeny of colubrine snakes in Southwestern Asia, with description of a new genus and species (Serpentes: Colubridae: Colubrinae). PeerJ 8: e9016. DOI: 10.7717/peerj.9016 Reference page. 

2021

• Burbrink, F.T., Pyron, R.A., Gehara, M., McKelvy, A.D. & Myers, E.A. 2021. The corrected taxonomic history of the North American ratsnakes (Pantherophis obsoletus complex). Herpetological Review 52(3): 537–547. Reference page. 
• Dubois, A., Ohler, A. & Pyron, R.A. 2021. New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians. Megataxa 5(1): 1–738. DOI: 10.11646/MEGATAXA.5.1.1 Reference page. 
• Hime, P.M., Lemmon, A.R., Lemmon, E.M., Prendini, E., Brown, J.M., Thomson, R.C., Kratovil, J.D., Noonan, B.P., Pyron, R.A., Peloso, P.L.V., Kortyna, M.L., Keogh, J.S., Donnellan, S.C., Mueller, R.L., Raxworthy, C.J., Kunte, K., Ron, S.R., Das, S., Gaitonde, N., Green, D.M., Labisko, J., Che, J. & Weisrock, D.W. 2021. Phylogenomics reveals ancient gene tree discordance in the amphibian tree of life. Systematic Biology 70(1): 49–66 DOI: 10.1093/sysbio/syaa034 Reference page. 
• Longrich, N.R., Suberbiola, X. P., Pyron, R.A., & Jalil, N.-E. 2021. The first duckbill dinosaur (Hadrosauridae: Lambeosaurinae) from Africa and the role of oceanic dispersal in dinosaur biogeography. Cretaceous Research 2020: 104678. DOI: 10.1016/j.cretres.2020.104678 Reference page. 

2022

• Pyron, R.A. & Beamer, D.A. 2022. A nomenclatural and taxonomic review of the salamanders (Urodela) from Holbrook’s North American Herpetology. Zootaxa 5188(6): 151–196. DOI: 10.11646/zootaxa.5188.6.1 Reference page. 
• Pyron, R.A. & Beamer, D.A. 2022. Nomenclatural solutions for diagnosing ‘cryptic’ species using molecular and morphological data facilitate a taxonomic revision of the Black-bellied Salamanders (Urodela, Desmognathus ‘quadramaculatus’) from the southern Appalachian Mountains. Bionomina 27(1): 1–43. DOI: 10.11646/bionomina.27.1.1 Reference page. 
• Pyron, R.A. & Beamer, D.A. 2022. Allocation of Salamandra auriculata Holbrook, 1838, with a new species of swamp-dwelling dusky salamander (Plethodontidae: Desmognathus) from the Atlantic Coastal Plain. Zootaxa 5188(6): 587–595. DOI: 10.11646/zootaxa.5188.6.6 Reference page. 
• Pyron, R.A. & Beamer, D.A. 2022. Systematics of the Ocoee Salamander (Plethodontidae: Desmognathus ocoee), with description of two new species from the southern Blue Ridge Mountains. Zootaxa 5190(2): 207–240. DOI: 10.11646/zootaxa.5190.2.3 Reference page. 
• Pyron, R.A., O’Connell, K.A., Lemmon, E.M., Lemmon, A.R. & Beamer, D.A. 2022. Candidate-species delimitation in Desmognathus salamanders reveals gene flow across lineage boundaries, confounding phylogenetic estimation and clarifying hybrid zones. Ecology and Evolution 12(2): e8574. DOI: 10.1002/ece3.8574 Reference page. 
• Pyron, R.A., Beamer, D.A., Holzheuser, C.R., Lemmon, E.M., Lemmon, A.R., Wynn, A.H. & O’Connell, K.A. 2022. Contextualizing enigmatic extinctions using genomic DNA from fluid-preserved museum specimens of Desmognathus salamanders. Conservation Genetics 23: 375–386. DOI: 10.1007/s10592-021-01424-4 Reference page. 
• Pyron, R.A., O’Connell, K.A., Lamb, J.Y. & Beamer, D.A. 2022. A new, narrowly endemic species of swamp-dwelling dusky salamander (Plethodontidae: Desmognathus) from the Gulf Coastal Plain of Mississippi and Alabama. Zootaxa 5133(1): 53–82. DOI: 10.11646/zootaxa.5133.1.3 Reference page. 

2023

• Pyron, R.A. & Beamer, D.A. 2023. A systematic revision of the Shovel-nosed Salamander (Plethodontidae: Desmognathus marmoratus), with re-description of the related D. aureatus and D. intermedius. Zootaxa 5270(2): 262–280. DOI: 10.11646/zootaxa.5270.2.5 Reference page. 
• Pyron, R.A. & Beamer, D.A. 2023. Systematic revision of the Spotted and Northern Dusky Salamanders (Plethodontidae: Desmognathus conanti and D. fuscus), with six new species from the eastern United States. Zootaxa 5311(4): 451–504. DOI: 10.11646/zootaxa.5311.4.1 Reference page. 
• Pyron, R.A., O’Connell, K.A., Duncan, S.C., Burbrink, F.T. & Beamer, D.A. 2023. Speciation Hypotheses from Phylogeographic Delimitation Yield an Integrative Taxonomy for Seal Salamanders (Desmognathus monticola). Systematic Biology 72(1): 179–197. DOI: 10.1093/sysbio/syac065 Reference page. 
• Ray, J.M., Sánchez-Martínez, P., Batista, A., Mulcahy, D.G., Sheehy, C.M., Smith, E.N., Pyron, R.A. & Arteaga, A. 2023. A new species of Dipsas (Serpentes, Dipsadidae) from central Panama. Zookeys 1145: 131–167. DOI: 10.3897/zookeys.1145.96616 Reference page. 

2024

• Arteaga, A., Pyron, R.A., Batista, A., Vieira, J., Meneses-Pelayo, E., Smith, E.N., Barrio Amorós, C.L., Koch, C., Agne, S., Valencia, J.H., Bustamante, L. & Harris, K.J. 2024. Systematic revision of the Eyelash Palm-Pitviper Bothriechis schlegelii (Serpentes, Viperidae), with the description of five new species and revalidation of three. Evolutionary Systematics 8(1): 15–64. DOI: 10.3897/evolsyst.8.114527 Reference page. 
• Burbrink, F.T., Myers, E.A. & Pyron, R.A. 2024. Understanding species limits through the formation of phylogeographic lineages. Ecology and Evolution 14(10): e70263. DOI: 10.1002/ece3.70263 Reference page. 
• Dalman, S.G., Loewen, M.A., Pyron, R.A., Jasinski, S.E., Malinzak, D.E., Lucas, S.G., Fiorillo, A.R., Currie, P.J. & Longrich, N.R. 2024. A giant tyrannosaur from the Campanian–Maastrichtian of southern North America and the evolution of tyrannosaurid gigantism. Scientific Reports 13(22124): 1–11. DOI: 10.1038/s41598-023-47011-0 Reference page.