Achyranthes sandwicensis

Achyranthes sandwicensis

Taxonavigation

Taxonavigation: Caryophyllales
Classification System: APG IV Superregnum: Eukaryota Cladus: Diaphoretickes Cladus: CAM Cladus: Archaeplastida Regnum: Plantae Cladus: Angiosperms Cladus: Eudicots Cladus: Core eudicots Ordo: Caryophyllales

Familia: Amaranthaceae s.l.
Cladus: Amaranthaceae s.str.
Subfamilia: Amaranthoideae
Genus: Achyranthes
Species: Achyranthes sandwicensis

Name

Achyranthes sandwicensis (A.Gray) Di Vincenzo, Berends., Wondafr. & Borsch, Taxon 74(1): 84 (2024).

Synonyms

• Basionym
  • Ptilotus sandwicensis A.Gray ex H.Mann, Proc. Amer. Acad. Arts 7: 200. (1867)
    • Lectotype (designated by Sherff 1951, Bot. Leafl. 4: 6.): U.S.A. Hawaii, Oahu, near the coast, J. Rémy 207'. GH GH00037133. Isolectotypes: BISH BISH1005572, F, NY 00341968.
    • Additional syntype: Hawai`i, U.S. Expl. Exped. s.n. (US).
• Homotypic
  • Psilotrichum sandwicense (A.Gray) Seem., Fl. Vit. 198. (1867)
  • Nototrichium sandwicense (A.Gray) Hillebr., Fl. Hawaiian Isl. [Hillebrand] 373. (1888)
• Heterotypic
  • Ptilotus sandwicensis var. kavaiensis A.Gray ex H.Mann, Proc. Amer. Acad. Arts 7: 200. (1867)
    • Psilotrichum sandwicense var. kauaiense (A.Gray ex H.Mann) Drake, Ill. fl. ins. pacif. 270. (1886)
    • Nototrichium sandwicense var. kauaiense (A.Gray ex H.Mann) Hillebr., Fl. Hawaiian Isl. 373. 1888.
      • Holotype: U.S.A. Hawaii, Coast of Kauai, 1838–1842, U.S. South Pacific Exploring Expedition under the command of Capt. Wilkes, U.S.N., s.n. US 00462185.
  • Nototrichium sandwicense var. longespicatum Hillebr., Fl. Hawaiian Isl. 373. (1888), (as "longe-spicatum")
    • Psilotrichum sandwicense var. longespicatum (Hillebr.) Drake, Ill. fl. ins. pacif. 270. (1886)
      • Lectotype (designated by Sherff 1951, Bot. Leafl. 4: 9.): Maui: West Maui. W. Hillebrand s.n.. B (destroyed).
      • Neotype (designated by H.St. John 1979, Phytologia 42: 27.): U.S.A. Hawaiian Islands, Moloka`i: Settlement Trail [Kalaupapa]. June 1912. C.N. Forbes 43.Mo. BISH BISH1005560. ⧼Isoneotypes⧽: BISH BISH1005558, BISH1005559 , NY, US US00462171.
      • Additional syntype: Moloka`i, Kalaupapa, Hillebrand s.n. See neotypification.
  • Nototrichium viride Hillebr., Fl. Hawaiian Isl. 373. (1888)
    • Psilotrichum viride (Hillebr.) Drake, Ill. Fl. Ins. Pacif. 270. (1892)
      • Holotype: U.S.A. Hawaii, Kaua`i, Hanapepe. 1864–1865. H. Mann & W.T. Brigham 590. CU, incorporated in BH in 1977. Isotypes: B [PD, fragment BISH], BISH BISH1005567, BISH1005568, GH 00037128, K K000814959 , NY, US 00102713.
  • Nototrichium fulvum Schinz, Vierteljahrsschr. Naturf. Ges. Zürich 76: 142. (1931)
    • Holotype: U.S.A. Hawaii: Kaua`i, Kaholuamanu, above Waimea, between forks of Waimea River [on exposed slope on ridge up to Kaholuamanu at 2500 ft]. 24-30 Sept. 1895. A.A. Heller 2831 pro parte. Z+ZT Z-000000320. Isotypes: BISH, MO No. 37457 (MO-216273), NY 00324551, US 00102707.
  • Nototrichium sandwicense var. decipiens Sherff, Bot. Leafl. 2: 2. (1950)
    • Holotype: U.S.A. Hawaiian Islands: Kauai: on ridge leading up to Kaholuamanu, between forks of Waimea River, on open exposed slope. 2500ft. 24 Sept. 1895. A.A. Heller 2831 pro parte. US 00102707 Isotypes: AC 00312958, BISH, MO No. 37457 (MO-216373), NY 00341951, Z.
  • Nototrichium sandwicense var. dubium Sherff, Sherff, Bot. Leafl. 2: 2. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Maui: southeastern East Maui, Manawainui, south slope of Haleakala, on side of a gulch. 3 March 1920. C.N. Forbes 1816 M. BISH BISH1005543. Isotypes: A 00037121, BISH BISH1005544, BISH1005545, K K000814958, L L0038796, PTBG PTBG 1000000022, US 00102708, 00516918.
  • Nototrichium sandwicense var. forbesii Sherff, Bot. Leafl. 2: 2. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Moloka`i: Mauna Loa. June 1912. C.N. Forbes 3.Mo. NY NY00341952. Isotypes: BISH BISH1005546, BISH1005547, BISH1005548, BISH1005549, K K000814957, MO No. 788105 (MO-247247), US (2).
  • Nototrichium sandwicense var. helleri Sherff, Bot. Leafl. 1: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Kauai: on ridge leading up to Kaholuamanu, between forks of Waimea River. 24 September 1895. A.A. Heller 2831 pro parte. F No. 429251 (V0047630F). Isotype: GH 00037122.
  • Nototrichium sandwicense var. kolekolense Sherff, Bot. Leafl. 1: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Molokai: on slopes of Puu Kolekole. 9 July 1912. C.N. Forbes 179.Mo. F No. 485227 (V0047631F). Isotypes: BISH BISH1005550, BISH1005551, MO No. 786771 (MO-216355).
  • Nototrichium sandwicense var. lanaiense Sherff, Bot. Leafl. 1: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Lanai: W end [of island], in dry forests. June 1913. C.N. Forbes 155.L. BISH BISH1005552. Isotypes: BISH BISH1005553, CAS 0007266, 0007267, US 00102711.
  • Nototrichium sandwicense var. lanceolatum Sherff, Bot. Leafl. 1: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Hawaii: Kona, Puuwaawaa. August 1917. J.F. Rock 12973. BISH BISH1005554. Isotype: BISH BISH1005555.
  • Nototrichium sandwicense var. latifolium Sherff, Bot. Leafl. 1: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Lanai: Maunalei Gorge. 30 July 1910. J.F. Rock & J.S. Hammond 8117. GH GH00037124. Isotype: BISH BISH1005556, NY 00341955.
  • Nototrichium sandwicense var. leptopodum O.Deg. & Sherff, in Sherff, Bot. Leafl. 1: 2. 1950.
    • Holotype: U.S.A. Hawaiian Islands, Maui: Nakaohu, on `a`a lava waste. 23 December 1948. O. Degener 19341. F No. 1449387 F0047632F. Isotypes: BISH BISH1005557, GH 00037125, P P00636905, US 00462159.
  • Nototrichium sandwicense var. macrophyllum Sherff, Bot. Leafl. 1: 2. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Hawaii: Kau, Hilea, between Hilea and Honuapo, Government Road, old `a`a laval flow. alt. 125 ft. 3 June 1927. G.W. Russ s.n.. BISH BISH1005561. Isotype: BISH BISH1005562.
  • Nototrichium sandwicense var. mauiense O.Deg. & Sherff, in Sherff, Bot. Leafl. 2: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Maui: from Papawai Point, toward Puu Anu through Manawainui Gulch, arid region near cliff. 12 July 1927. O. Degener & H. Wiebke 3341. NY NY00341956.
  • Nototrichium sandwicense var. olokeleanum Sherff, Bot. Leafl. 2: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Kauai: Olokele Canyon. 1500ft. September 1909. J.F. Rock 5422. GH GH00037126. Isotypes: BISH BISH1005536, US 00102712.
  • Nototrichium sandwicense var. pulchellum Sherff, Bot. Leafl. 2: 3. (1950)
    • Holotype: U.S.A. Hawaiian Islands, Western Molokai, Mahana. February 1910. J.F. Rock s.n.. GH GH00037127. Isotypes: BISH BISH1005565, BISH1005566, NY 00341958.
  • Nototrichium sandwicense var. subcordatum O.Deg. & Sherff, in Sherff, Bot. Leafl. 1: 3. 1950.
    • Holotype: U.S.A. Hawaiian Islands, Hawaii, ca. 17 mi from Kohala toward Waimea. 13 August 1926. O. Degener 9873. NY 00341959.
  • Nototrichium sandwicense var. syringifolium Sherff, Bot. Leafl. 1: 2. (1950)
    • Holotype: U.S.A. Hawaiian Islands: 1851–1871, W. Hillebrand s.n.. K K000814960.
  • Nototrichium viride var. oblongifolium Sherff, Bot. Leafl. 2: 3. (1950)
    • Holotype: U.S.A. Kaua‘i: Waimea, rocky places of mountains Island Oahu, Hawaian Islands. March 1910. U. Faurie 1070. G-DEL. Isotype: BISH BISH1005570.
  • Nototrichium viride var. subtruncatum Sherff, Bot. Leafl. 2: 4. (1950)
    • Holotype: U.S.A. O‘ahu: Kawaihapae [Kawaihapai]. March 1932. G.W. Russ s.n.. BISH BISH1005571.
  • Nototrichium sandwicense var. pulchelloides O.Deg. & Sherff, in Sherff, Bot. Leafl. 3: 2. 1951.
    • Holotype: U.S.A. Hawaiian Islands, West Molokai, near Waiahewahewa Gulch, on cliff, in last relic of arid forest. 18 April 1928. O. Degener 9869. NY 00341957.
  • Nototrichium sandwicense var. niihauense H.St.John, Pacific Sci. 13: 166. 1959.
    • Holotype: U.S.A. Hawaii. Ni‘ihau: first valley W of Ka`ali Cliff, top of steep basalt talus. 100ft. 16 August 1947. H.St. John 22830. BISH BISH1005563. Isotypes: BISH BISH1005564, K K000814961, L 2D-barcode L.3725009, CANB CBG8100594, P P00636906.
  • Nototrichium viride var. kalaupapae H.St.John, Phytologia 45: 295. (1980)
    • Holotype: U.S.A. Hawaiian Islands, Molokai Island: Kalaupapa, Uau Crater [Pu`u Uao], inside crater. 8 March 1974. B. Harrison & W. Gagné s.n.: BISH BISH1005569.

Distribution

Native distribution areas:
Pacific North-Central Pacific Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Kahoolawe Isl., Hawaii Isl.) References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition

References

Primary references

• Di Vincenzo, V., Berendsohn, W., Wondafrash, M. & Borsch, T. 2025. Phylogenetics and morphological character evolution in the achyranthoid clade (Amaranthaceae): Evidence to re-circumscribe the genera Achyranthes and Cyathula and to resurrect a third species of the former genus Sericocomopsis in East Africa. Taxon 74(1): 66–100. DOI: 10.1002/tax.13284 Reference page. 
• Drake del Castillo, E. 1886 [–1892]. Illustrationes Florae Insularum Maris Pacifici. Parisiis: G. Masson. BHL Reference page. 
• Hillebrand, W. (†). 1888. Flora of the Hawaiian Islands: a description of their phanerogams and vascular cryptogams. London: Williams & Norgate; New York: B. Westermann & Co.; Heidelberg: Carl Winter. BHL Reference page. 
• Mann, H. 1867. Enumeration of Hawaiian plants. Proceedings of the American Academy of Arts and Sciences 7: 143–222. BHL Reference page. 
• Sherff, E.E. 1950. Botanical Leaflets, a Series of Studies in the Systematic Botany of Miscellaneous Dicotyledonous Plants. Chicago. Vol. 1. Reference page. 

Links

• Govaerts, R. et al. 2025. Achyranthes sandwicensis in Kew Science Plants of the World Online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2025 Nov. 26. Reference page. 
• International Plant Names Index. 2025. Achyranthes sandwicensis. Published online. Accessed: Nov. 26 2025. Reference page.

Vernacular names

• English: Hawai'i rockwort
• Hawaiʻi: Kuluʻī

For more multimedia, look at Nototrichium sandwicense on Wikimedia Commons.