Munetoshi Maruyama

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Munetoshi Maruyama, Japanese entomologist.

  • Department of Zoology, National Science Museum (Nat. Hist.), 3-23-1 Hyakunin-chô, Shinjuku-ku, Tôkyô-to, 169-0073 Japan.
  • E-mail:
  • The Kyushu University Museum, Fukuoka, 812-8581 Japan.
  • E-mail:

Taxon names authored

(List may be incomplete)



(List may be incomplete)




  • Naomi, S & Maruyama, M. 1998. A revision of the genus Sepedophilus Gistel (Coleoptera, Staphylinidae, Tachyporinae) from Japan: Species group of S. pedicularius. Japanese journal of systematic entomology, 4(1): 51–75.
  • Naomi, S & Maruyama, M. 1998. Three new species of the genus Sepedophilus Gistel (Coleoptera, Staphylinidae, Tachyporinae) from Taiwan. Natural History Research, 5(1): 43–51.


  • Ahn, K-J., Maruyama, M. & Ôhara, M. 1999. The intertidal beetle Amblopusa magna Zerche (Coleoptera, Staphylinidae, Aleocharinae), new to Hokkaido, Japan, and Kuril Archipelago, Russia. Elytra, Tokyo 27(2): 641–642.
  • Maruyama, M. & Y. Shibata 1999. Notes on the myrmecophilous rove beetle Ophryomedon marginatus Naomi (Coleoptera, Staphylinidae, Paederinae). Elytra, Tokyo, 27(2): 518.
  • Maruyama, M & T. Kishimoto, 1999. A revision of the Japanese species of the genus Tetrabothrus (Coleoptera, Staphylinidae, Aleocharinae). Elytra, Tokyo, 27(2): 503–517.
  • Hoshina, H. & Maruyama, M. 1999. An additional new species to the Scaphidium fauna (Coleoptera, Staphylinidae, Scaphidiinae) of Ryukyus, Japan. Elytra, Tokyo, 27(2): 479–484.




  • Hoshina, H & Maruyama, M. 2001. New record of the genus Agathidium (Coleoptera: Leiodidae) from Rishiri Is., Hokkaido, Japan. Bulletin of the Institute of Environmental Science and Technology, Yokohama National University, 27(1): 157–159.
  • Maruyama, M. 2001. Aleochara (Xenochara) fumata Gravenhorst (Coleoptera, Staphylinidae, Aleocharinae), new to Japan. Elytra, Tokyo, 29(2): 486.




  • Maruyama, M. & S.H. Yek, R. Hashim, & F. Ito 2003. A new myrmecophilous species of Drusilla (Coleoptera, Staphylinidae, Aleocharinae) from Peninsular Malaysia, a possible Batesian mimic associated with Crematogaster inflata (Hymenoptera, Formicidae, Myrmicinae). Japanese journal of systematic entomology, 9(2): 267–275.
  • Maruyama, M. & P. Hlavac 2003. A new genus and species of the tribe Aleocharini (Coleoptera, Staphylinidae, Aleocharinae) from Morocco, associated with Aphaenogaster (Hymenoptera, Formicidae, Myrmicinae). Annales de la Societe Entomologique de France, 39(3): 225–228.(PDF)
  • Ahn, K-J., Maruyama, M. & Jeon, M-J. 2003. Redescription of Brachypronomaea esakii and its systematic position (Coleoptera, Staphylinidae, Aleocharinae). Journal of the Kansas Entomological Society 76(4): 622–629. JSTOR Reference page
  • Maruyama, M. & Ahn, K-J. 2003. New records of Zyras (Coleoptera, Staphylinidae, Aleocharinae) from Korea. Elytra, Tokyo 31(1): 262.
  • Maruyama, M. & T. Miyake 2003. Additions to the subgenus Ectolabrus of the genus Euryusa (Coleoptera, Staphylinidae, Aleocharinae), with description of a new species and revised key to species. Entomological review of Japan, 58(1): 71–78.
  • Sugaya, H. & Maruyama, M. 2003. Additional records of Triartiger klapperichorum from Taiwan, collected by the use of flight interception traps (Coleoptera, Staphylinidae, Pselaphinae). Entomological review of Japan 58(1): 11–12.
  • Sugaya, H. & Maruyama, M. 2003. A host record of the myrmecophilous Micrelytriger loebli (Coleoptera, Staphylinidae, Pselaphinae), with additional locality records in Taiwan. Entomological review of Japan, 58(1): 9–10.
  • Sugaya, H. & Maruyama, M. 2003. Biological notes on Tribasodites picticornis, with new locality records in the Ryukyus, Japan (Coleoptera, Staphylinidae, Pselaphinae). Entomological review of Japan, 58(1): 7–8.


  • Maruyama, M. 2004. Two new species of the Lomechusini (Coleoptera: Staphylinidae: Aleocharinae) from Laos. Entomological review of Japan 59(1): 87–97.
  • Maruyama, M. & T. Hironaga 2004. Microdon katsurai, a new species of myrmecophilous hoverfly (Diptera, Syrphidae) from Japan, associated with Polyrhachis lamellidens (Hymenoptera, Formicidae). Bulletin of National Science Museum, Tokyo, Ser. A, Ser. A, Zoology 30(4): 173–179.
  • Maruyama, M. 2004. Chasmatodera grata, a new species of termitophilous pleasing fungus beetle from Thailand (Coleoptera, Erotylidae, Tritominae). Japanese journal of systematic entomology, 10(2): 63–68.
  • Maruyama, M. 2004. Symbiotic host of the myrmecophilous histerid beetle, Hetaerius optatus (Coleoptera, Histeridae, Hetaeriinae). Elytra, Tokyo 32(2): 326.
  • Maruyama, M. & H. Sugaya 2004. A new myrmecophilous species of Batraxis (Coleoptera, Staphylinidae, Pselaphinae), associated with Lasius (Chthonolasius) sp. (Hymenoptera, Formicidae, Formicinae) from Hokkaido, Japan. Elytra, Tokyo 32(2): 321–325.
  • Maruyama, M. & Klimaszewski, J. 2004. A new species of the myrmecophilous genus Goniusa (Coleoptera, Staphylinidae, Aleocharinae) from Canada. Elytra, Tokyo 32(2): 315–320.
  • Maruyama, M. & A. Smetana 2004. Myrmecophilous species of the Quedius hirticornis Sharp, 1889 complex (Coleoptera, Staphylinidae, Staphylininae). Elytra, Tokyo 32(2): 299–313.
  • Maruyama, M. 2004. A permanent slide under a specimen. Elytra, Tokyo 32(2): 276.
  • Maruyama, M. 2004. Drusilla keiichii (Coleoptera: Staphylinidae: Aleocharinae), a new termitophile associated with Hospitalitermes (Isoptera: Termitidae) from Thailand. Entomological review of Japan 59 (2): 249–254.
  • Maruyama, M. & Klimaszewski, J. 2004. A new genus and species of the myrmecophilous Athetini, Paragoniusa myrmicae (Coleoptera: Staphylinidae: Aleocharinae) from Canada. Entomological review of Japan 59(2): 241–248.
  • Maruyama, M. 2004. Redescription of Creochara (Coleoptera, Staphylinidae, Aleocharinae, Aleocharini) and its systematic position. Canadian entomologist 136: 621–637.
  • Maruyama, M. 2004. On the systematic position of Amaurodera wallacei Kistner & McNairn (Coleoptera, Staphylinidae, Aleocharinae). Koleopterologische rundschau 74: 220.
  • Maruyama, M. 2004. Taxonomy of the myrmecophilous genus Myrmecopella (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of seven new species. Koleopterologische rundschau 74: 201–219.
  • Hlaváč, P. & Maruyama, M. 2004: A new genus and species of the myrmecophilous tribe Sahlbergiini from Malaysia (Coleoptera: Staphylinidae: Aleocharinae). Koleopterologische rundschau 74: 191–199. Full article (PDF)
  • Hironaga, T. & Maruyama, M. 2004. The myrmecophilous hoverfly genus Microdon (Diptera, Syrphidae, Microdontinae) of Hokkaido, Japan, with descriptions of four new species. Bulletin of National Science Museum, Tokyo, Ser. A, Zoology 30(2): 87–103.
  • Maruyama, M., M. Ohara, A. Woro, & S. Hartini 2004. A new species of Delibius (Coleoptera, Staphylinidae, Aleocharinae, Pygostenini) from Jawa Barat, Indonesia. Japanese journal of systematic entomology 10(1): 63–68.
  • Maruyama, M. 2004. Two new species of Lomechusini (Coleoptera, Staphylinidae, Aleocharinae) from Laos. Entomological review of Japan 59(1): 87–97.
  • Maruyama, M. 2004. Additional records of Philetaerius elegans (Coleoptera, Staphylinidae, Staphylininae). Elytra, Tokyo 32(1): 114.
  • Maruyama, M. & P. Hlavac 2004. Two new species of the myrmecophilous genus Lomechusoides (Coleoptera, Staphylinidae, Aleocharinae, Lomechusini) from Sichuan, China. Elytra, Tokyo 32(1): 105–113.
  • Maruyama, M. 2004. Four new species of the genus Myrmecophilus (Orthoptera, Myrmecophilidae) from Japan. Bulletin of National Science Museum, Tokyo, Ser. A, Zoology 30(1): 37–44.


  • Maruyama, M. 2005. Some additional or new records of Paussinae (Coleoptera, Carabidae) from Asia. Elytra, Tokyo, 33(2): 648.
  • Maruyama, M. 2005. A new record of Margarinotus maruyamai (Coleoptera, Histeridae) from the Russian Far East. Elytra, Tokyo, 33(2): 616.
  • Maruyama, M. 2005. A new synonymy in the subgenus Dendrolasius of the genus Lasius (Hymenoptera, Formicidae, Formicinae). Bulletin of National Science Museum, Tokyo, Ser. A, Zoology, 31(3): 115–117.
  • Taniguchi, Maruyama, M., T. Ichikawa, & F. Ito 2005. A case of Batesian mimicry between a myrmecophilous staphylinid beetle, Pella comes, and its host ant, Lasius (Dendrolasius) spathepus: an experiment using the Japanese treefrog, Hyla japonica as a real predator. Insectes Sociaux, 52(4): 320–322.
  • Klimaszewski, J, G. Pelletier, Maruyama, M., & P. Hlavac 2005. Canadian species of the Zyras group of genera and review of the types from America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Revue suisse de zoologie, 122(3): 703–733.


  • Shibata, Y., Maruyama, M., & M. Ohara 2006. The subfamily Staphylininae (Coleoptera, Staphylinidae) fin the Kuril Archipelago. Biodiversity and Biogeography of the Kuril Islands and Sakhalin, 2: 151–159.
  • Maruyama, M. & J. Klimaszewski 2006. Notes on myrmecophilous aleocharines (Insecta, Coleoptera, Staphylinidae) from Canada, with a description of a new species of Myrmoecia. Bulletin of National Science Museum, Tokyo, Ser. A, Zoology, 32(3): 125–131.
  • Maruyama, M. 2006. Turcizyras assingi, a new genus and species of the tribe Lomechusini (Staphylinidae, Aleocharinae) from Turkey. Bulletin of National Science Museum, Tokyo, Ser. A, Zoology 32(2): 47–51.
  • Maruyama, M. 2006. Revision of the Palearctic species of the myrmecophilous genus Pella (Coleoptera, Staphylinidae, Aleocharinae). National Science Museum Monographs: 32. 1–207.


  • Hamaguti K, Matsumoto T, Maruyama M, Hashimoto Y, Yamane S, Itioka T. 2007: Isolation and characterization of eight microsatellite loci in two morphotypes of the Southeast Asian army ant, Aenictus laeviceps. Molecular Ecology Notes 7: 984–986.


  • Maruyama, M. 2008. Giraffaenictus eguchii (Coleoptera, Staphylinidae, Aleocharinae), a new genus and species of fully myrmecoid myrmecophile from a colony of Aenictus binghami (Hymenoptera, Formicidae, Aenictinae) in Vietnam. Esakia 48: 51–56. PDF Reference page
  • Maruyama, M., Y. Katayama, W. Sakchoowong & S. Nomura 2008. First record of Lebioderus (Coleoptera, Carabidae, Paussinae) from the Indochinese Peninsula, with description of a new species. Esakia, 48: 47–49.
  • Komatsu T., Maruyama, M., S. Ueda & T. Itinio 2008. mtDNA phylogeny of Japanese ant crickets (Orthoptera: Myrmecophilidae) : Diversification in host specificity and habitat use. Sociobiology, 52: 553–565.
  • Maruyama, M., H. R. Disney & R. Hashim 2008. Three new species of legless, wingless scuttle flies (Diptera: Phoridae) associated with army ants (Hymenoptera: Formicidae) in Malaysia. Sociobiology, 52: 485–496.
  • Maruyama M., F. M Steiner, C. Stauffer, T. Akino, R. H Crozier, & B. C Schlick-Steiner 2008. A DNA and morphology based phylogenetic framework of the ant genus Lasius with hypotheses for the evolution of social parasitism and fungiculture. BMC Evolutionary Biology, 2008, 8: 237.
  • Maruyama M.. & Disney R.H.L. 2008. Scuttle flies associated with Old World army ants in Malaysia (Diptera: Phoridae; Hymenoptera, Formicidae, Dorylinae). Sociobiology, 51: 65–71.
  • Komatsu, T. & Maruyama, M. 2008. Symbiotic host of Triartiger reductus Nomura (Coleoptera, Staphylinidae, Pselaphinae, Clavigeritae) in the Island of Tsushima, Japan. Elytra, Tokyo, 36: 227–228.
  • Maruyama, M., L. Brian Patrick & V. Gusarov 2008. First record of the genus Myrmedonota Cameron (Coleoptera, Staphylinidae) from North America, with descriptions of two new species. Zootaxa 1716: 35–43.
  • Maruyama, M., Klimaszewski, J. & Gusarov, V.I. 2008. Osakatheta yasukoae, a new intertidal genus and species of athetine rove beetles (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 1683: 39–50. Abstract & excerpt. Reference page


  • Maruyama, M. 2009. On the myrmecophilous genus Losiusa Seevers, 1978 (Coleoptera, Staphylinidae, Aleocharinae). Esakia, 49: 111–116.
  • Maruyama, M. 2009. A new species of the genus Merismoderus (Coleoptera, Carabidae, Paussinae) from Laos, with a revised key of the genus and a new combination. Esakia, (49): 107–109.
  • Maruyama, M. 2009. Rhinocerotopsis nakasei (Coleoptera, Scarabaeidae, Aphodiinae), a new genus and species of Stereomerini from Peninsular Malaysia. Esakia, 49: 103–106.
  • Komatsu, T., Maruyama, M., & T. Itino 2009. Behavioral differences between two ant cricket species in Nansei Islands: host-specialist versus host-generalist. Insectes sociaux, 56: 389–396.
  • Maruyama, M. & Hayashi, M. 2009. Description of the intertidal aleocharine Halorhadinus sawadai sp.n. from Japan, with notes on the genus Halorhadinus Sawada, 1971 (Coleoptera: Staphylinidae). Koleopterologische rundschau 79: 71–82. Reference page
  • Yamamoto, S. & Maruyama, M. 2009. Description of Aleochara (Maseochara) hiranoi sp.n. from Japan (Coleoptera: Staphylinidae: Aleocharinae). Koleopterologische rundschau, 79: 65–70.
  • Maruyama M., T. Akino, R. Hashim & T. Komatsu 2009. Behavior and cuticular hydrocarbons of myrmecophilous insects (Tysanura; Coleoptera: Staphylinidae; Diptera: Phoridae) associated with Asian Aenictus army ants (Hymenoptera; Formicidae). Sociobiology, 54: 19–35.


  • von Beeren, C., Maruyama, M., Rosli, H., Witte, V. 2010: Differential host defense against multiple parasites in ants. Evolutionary Ecology, 25: 259–276.
  • Maruyama, M., Beeren, C. von & Hashim, R. 2010. Aleocharine rove beetles (Coleoptera, Staphylinidae) associated with Leptogenys Roger, 1861 (Hymenoptera, Formicidae) I. Review of three genera associated with L. distinguenda (Emery, 1887) and L. mutabilis (Smith, 1861). ZooKeys 59: 47–60. DOI: 10.3897/zookeys.59.510 Reference page
  • Maruyama, M., Beeren, C. von & Witte, V. 2010. Aleocharine rove beetles (Coleoptera, Staphylinidae) associated with Leptogenys Roger, 1861 (Hymenoptera, Formicidae) II. Two new genera and two new species associated with L. borneensis Wheeler, 1919. ZooKeys 59: 61–72. DOI: 10.3897/zookeys.59.511 Reference page
  • Komatsu, T., Maruyama, M., Itino, T. 2010: Differences in host specificity and behavior of two ant cricket species (Orthoptera: Myrmecophilidae) in Honshu, Japan. Journal of Entomological Science, 45: 1–12.
  • Kanao, T., Maruyama, M. & Sakchoowong, W. 2010. Two new species of Aleocharinae (Coleoptera, Staphylinidae) found in fungus gardens of Odontotermes termites (Isoptera, Termitidae, Macrotermitinae) in Khao Yai National Park, Thailand. ZooKeys 49: 77–86. DOI: 10.3897/zookeys.49.468 Reference page
  • Ballerio, A., Maruyama, M. 2010. The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae). In: Ratcliffe B, Krell F-T (Eds) Current advances in Scarabaeoidea research. ZooKeys, 34: 77–104.
  • Maruyama, M. 2010. A new genus and species of myrmecophilous aphodiine beetle (Coleoptera, Scarabaeidae) inhabiting the myrmecophytic epiphyte Platycerium sp. (Polypodiaceae) in the Bornean rainforest canopy. In: Ratcliffe B., Krell F.-T. (Eds) Current advances in Scarabaeoidea research. ZooKeys, 34: 49–54.




  • Maruyama, M. 2012. Termitotrox cupido sp. n. (Coleoptera, Scarabaeidae), a new termitophilous scarab species from the Indo-Chinese subregion, associated with Hypotermes termites. ZooKeys, 254: 89–97.
  • Yamauchi, T., Takano, A., Maruyama, M., Kawabata, H. 2012. Human Infestation by Amblyomma testudinarium (Acari: Ixodidae) in Malay Peninsula, Malaysia. Journal of the Acarological Society of Japan, 21: 143–148.
  • Maruyama, M. 2012. A new genus and species of flightless, microphthalmic Corythoderini (Coleoptera: Scarabaeidae: Aphodiinae) from Cambodia, associated with Macrotermes termites. Zootaxa, 3555: 83–88.
  • Yamamoto, S., Maruyama, M. 2012. Revision of the seashore-dwelling subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan. Zootaxa, 3517: 1–52.
  • Maruyama, M., Kanao, T., Iwata, R. 2012. Discovery of two aleocharine staphylinid species (Coleoptera) associated with Coptotermes formosanus (Isoptera: Rhinotermitidae) from Central Japan, with a review of the possible natural distribution of C. formosanus in Japan and surrounding countries. Sociobiology, 59: 605–616.
  • Maruyama, M., Nomura, S., Sakchoowong, W. 2012. A new species of the genus Euplatyrhopalus (Coleoptera, Carabidae, Paussinae) from Thailand. Esakia, 52: 5–7.
  • Sutou, M., Maruyama, M., Komatsu, T., Kanao, T. 2012. Discovery of a remarkable new species of black fungus gnat (Diptera, Sciaridae) from termite nests in Malaysia. Journal of Natural History, 46(15–16): 969–978.
  • Kanao, T., Eldredge, K.T. & Maruyama, M. 2012. Two new genera and species of the termite symbiont lineage Termitohospitini (Coleoptera, Staphylinidae, Aleocharinae) from Bolivia and peninsular Malaysia. ZooKeys 254: 67–87. DOI: 10.3897/zookeys.254.4043 Reference page




















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