List of Tardigrada

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List of Tardigrada taxa, based on: Degma, P., Bertolani, R., Guidetti, R., 2012. Actual checklist of Tardigrada species (2009-2012, Ver. 20: 17–01-2012). [1]

Phylum: Tardigrada Spallanzani, 1777 (1354 Taxa, subdivided in: 3 classes, 5 ordines, 4 superfamiliae, 24 familiae, 12 subfamiliae, 113 genera, 4 subgenera, 1117 species, 71 subspecies)

Classis: Eutardigrada Marcus, 1927
Ordo: Apochela Schuster, Nelson, Grigarick and Christenberry, 1980
Familia: Milnesiidae Ramazzotti, 1962
Genus: Bergtrollus Dastych, 2011
Species: Bergtrollus dzimbowski Dastych, 2011
Genus: Limmenius Horning, Schuster and Grigarick, 1978
Species: Limmenius porcellus Horning, Schuster and Grigarick, 1978
Genus: Milnesioides Claxton, 1999
Species: Milnesioides exsertum Claxton, 1999
Genus: Milnesium Doyère, 1840
Species: Milnesium alabamae Wallendorf & Miller, 2009
Species: Milnesium almatyense Tumanov, 2006
Species: Milnesium antarcticum Tumanov, 2006
Species: Milnesium asiaticum Tumanov, 2006
Species: Milnesium brachyungue Binda and Pilato, 1991
Species: Milnesium dujiangensis Yang, 2003
Species: Milnesium eurystomum Maucci, 1991
Species: Milnesium granulatum Ramazzotti, 1962 <Amended and elevated to the species level by
Species: Milnesium jacobi Meyer & Hinton, 2010
Species: Milnesium katarzynae Kaczmarek, Michalczyk and Beasley, 2004
Species: Milnesium krzysztofi Kaczmarek & Michalczyk, 2007
Species: Milnesium longiungue Tumanov, 2006
Species: Milnesium minutum Pilato & Lisi, 2016
Species: Milnesium reductum Tumanov, 2006
Species: Milnesium reticulatum Pilato, Binda and Lisi, 2002
Species: Milnesium sandrae Pilato & Lisi, 2016
Species: Milnesium swolenskyi Bertolani and Grimaldi, 2000
Species: Milnesium tardigradum Doyère, 1840
Subspecies: Milnesium tardigradum tardigradum Doyère, 1840
Subspecies: Milnesium tardigradum trispinosum Rahm, 1931
Species: Milnesium tetralamellatum Pilato and Binda, 1991
Species: Milnesium zsalakoae Meyer & Hinton, 2010
Ordo: Parachela Schuster, Nelson, Grigarick and Christenberry, 1980
Superfamilia: Eohypsibioidea Bertolani & Kristensen, 1987 in Marley et al. 2011
Familia: Eohypsibiidae Bertolani & Kristensen, 1987
Synonym: Amphibolidae Bertolani, 1981
Genus: Austeruseus Trygvadóttir & Kristensen 2011
Species: Austeruseus balduri Trygvadóttir & Kristensen 2011
Species: Austeruseus faeroensis Trygvadóttir & Kristensen 2011
Species: Austeruseus rokuri Trygvadóttir & Kristensen 2011
Genus: Bertolanius Özdikmen, 2008
Synonym: Amphibolus Bertolani, 1981 in Özdikmen 2008
Species: Bertolanius mahunkai Iharos, 1971
Species: Bertolanius markevichi Biserov, 1992
Species: Bertolanius nebulosus Dastych, 1983
Species: Bertolanius portucalensis Fontoura, Pilato, Lisi & Morais, 2009
Species: Bertolanius smreczynskii Wêglarska, 1970


Species: Bertolanius volubilis Maucci, 1975


Species: Bertolanius weglarskae Dastych, 1972


Genus: Eohypsibius Kristensen, 1982
Species: Eohypsibius nadjae Kristensen, 1982
Species: Eohypsibius terrestris Ito, 1988
Superfamilia: Hypsibioidea Pilato, 1969 in Marley et al. 2011
Familia: Calohypsibiidae Pilato, 1969
Genus: Calohypsibius Thulin, 1928
Species: Calohypsibius maliki Michalczyk and Kaczmarek, 2005
Species: Calohypsibius ornatus Richters, 1900
Synonym: Hypsibius (Calohypsibius) armatus Bartoš, 1938
Synonym: Hypsibius (Calohypsibius) intermedius Mihelcic, 1939
Species: Calohypsibius placophorus da Cunha, 1943
Species: Calohypsibius schusteri Nelson and McGlothlin, 1996
Species: Calohypsibius verrucosus Richters, 1900
Synonym: Calohypsibius scabrosus Thulin, 1928


Genus: Haplohexapodibius Pilato and Beasley, 1987
Species: Haplohexapodibius seductor Pilato and Beasley, 1987
Synonym: Hexapodibius beasleyi May, 1948
Genus: Haplomacrobiotus May, 1948
Species: Haplomacrobiotus hermosillensis May, 1948
Species: Haplomacrobiotus utahensis Pilato and Beasley, 2005
Genus: Hexapodibius Pilato, 1969
Species: Hexapodibius bindae Pilato, 1982
Species: Hexapodibius boothi Dastych, 1994
Species: Hexapodibius christenberryae Pilato and Binda, 2003
Species: Hexapodibius micronyx Pilato, 1969
Species: Hexapodibius pseudomicronyx Robotti, 1972
Species: Hexapodibius reginae Vargha, 1995
Genus: Parhexapodibius Pilato, 1969
Species: Parhexapodibius bactrianus Biserov, 1999
Species: Parhexapodibius castrii Ramazzotti, 1964


Species: Parhexapodibius lagrecai Binda and Pilato, 1969


Species: Parhexapodibius pilatoi Bernard, 1977
Species: Parhexapodibius ramazzottii Manicardi & Bertolani, 1987
Familia: Hypsibiidae Pilato, 1969
Subfamilia: Diphasconinae Dastych, 1992
Genus: Bindius Pilato, 2009
Species: Bindius triquetrus Pilato, 2009
Genus: Diphascon Plate, 1888
Subgenus: Diphascon (Adropion) Pilato, 1987
Species: Diphascon (Adropion) arduifrons Thulin, 1928
Species: Diphascon (Adropion) behanae Dastych, 1987
Species: Diphascon (Adropion) belgicae Richters, 1911
Synonym: Fujiscon diphasconiellum Ito, 1991
Species: Diphascon (Adropion) carolae Binda & Pilato, 1969
Species: Diphascon (Adropion) clavatum Bartos, 1935
Species: Diphascon (Adropion) gordonense Pilato, Claxton and Horning, 1991
Species: Diphascon (Adropion) greveni Dastych, 1984
Species: Diphascon (Adropion) linzhiensis X. Li, 2007
Species: Diphascon (Adropion) maucci Dastych and McInnes, 1996
Species: Diphascon (Adropion) modestum Binda, Pilato & Dastych, 1984
Species: Diphascon (Adropion) montigenum Pilato & Dastych, 1979
Species: Diphascon (Adropion) onorei Pilato, Binda, Napolitano and Moncada, 2002
Species: Diphascon (Adropion) prorsirostre Thulin, 1928
Species: Diphascon (Adropion) scoticum Murray, 1905
Subspecies: Diphascon (Adropion) scoticum scoticum Murray, 1905
Synonym: Diphascon crozetense Richters, 1907
Subspecies: Diphascon (Adropion) scoticum ommatophorum Thulin, 1911
Subspecies: Diphascon (Adropion) scoticum qinlingensis X. Li & Liu, 2005
Species: Diphascon (Adropion) tricuspidatum Binda and Pilato, 2000
Species: Diphascon (Adropion) triodon Maucci, 1996
Subgenus: Diphascon (Diphascon) Plate, 1889
Species: Diphascon (Diphascon) aculeatum Maucci, 1952
Species: Diphascon (Diphascon) alpinum Murray, 1906
Species: Diphascon (Diphascon) australianum Binda and Pilato, 1998
Species: Diphascon (Diphascon) bidropion Ito, 1995
Species: Diphascon (Diphascon) birklehofi Schuster, 1999
Species: Diphascon (Diphascon) bisbullatum Iharos, 1964
Species: Diphascon (Diphascon) boreale Biserov, 1996
Species: Diphascon (Diphascon) brevipes Marcus, 1936
Species: Diphascon (Diphascon) bullatum Murray, 1905
Species: Diphascon (Diphascon) burti Nelson, 1991
Species: Diphascon (Diphascon) chilenense Plate, 1888
Species: Diphascon (Diphascon) claxtonae Pilato and Binda, 1998
Species: Diphascon (Diphascon) dastychi Pilato and Binda, 1999
Species: Diphascon (Diphascon) dolmiticum Pilato and Bertolani, 2005
Species: Diphascon (Diphascon) elongatum Mihelcic, 1959
Species: Diphascon (Diphascon) faialense Fontoura & Pilato, 2007
Species: Diphascon (Diphascon) gerdae Mihelcic, 1951
Species: Diphascon (Diphascon) granifer Greven, 1972
Species: Diphascon (Diphascon) halapiense Iharos, 1964
Species: Diphascon (Diphascon) higginsi Binda, 1971
Species: Diphascon (Diphascon) humicus Bertolani, Guidetti & Rebecchi, 1994
Species: Diphascon (Diphascon) hydrophilum Pilato, Binda, Bertolani and Lisi, 2005
Species: Diphascon (Diphascon) iharosi Vargha, 1995
Species: Diphascon (Diphascon) iltisi Schuster & Grigarick, 1965
Species: Diphascon (Diphascon) langhovdense Sudzuki, 1964
Species: Diphascon (Diphascon) latipes Mihelcic, 1955
Species: Diphascon (Diphascon) mirabilis Dastych, 1984
Species: Diphascon (Diphascon) mitrense Pilato, Binda and Quartieri, 1999
Species: Diphascon (Diphascon) nelsonae Pilato, Binda, Bertolani and Lisi, 2005
Species: Diphascon (Diphascon) nobilei Binda, 1969
Species: Diphascon (Diphascon) nodulosum Ramazzotti, 1957
Species: Diphascon (Diphascon) nonbullatum Mihelcic, 1951
Species: Diphascon (Diphascon) oculatum Murray, 1906
Subspecies: Diphascon (Diphascon) oculatum alpinum Mihelcic, 1964
Subspecies: Diphascon (Diphascon) oculatum oculatum Murray, 1906
Subspecies: Diphascon (Diphascon) oculatum canadense Murray, 1910
Synonym: Diphascon (Diphascon) oculatum vancouverense Thulin, 1911
Species: Diphascon (Diphascon) ongulense Morikawa, 1962
Species: Diphascon (Diphascon) opisthoglyptum Maucci, 1987
Species: Diphascon (Diphascon) patanei Binda & Pilato, 1971
Species: Diphascon (Diphascon) pingue Marcus, 1936
Subspecies: Diphascon (Diphascon) pingue pingue Marcus, 1936
Subspecies: Diphascon (Diphascon) pingue brunsvicense Argue, 1972
Species: Diphascon (Diphascon) pinguiforme Pilato & Binda, 1998
Species: Diphascon (Diphascon) platyungue Pilato, Binda, Bertolani and Lisi, 2005
Species: Diphascon (Diphascon) polare Pilato and Binda, 1999
Species: Diphascon (Diphascon) puniceum Jennings, 1976
Species: Diphascon (Diphascon) ramazzottii Robotti, 1970
Species: Diphascon (Diphascon) recamieri Richters, 1911
Species: Diphascon (Diphascon) rugocaudatum Rodriguez Roda, 1952
Species: Diphascon (Diphascon) rugosum Bartos, 1935
Species: Diphascon (Diphascon) sanae Dastych, Ryan and Watkins, 1990
Species: Diphascon (Diphascon) secchii Bertolani & Rebecchi, 1996
Species: Diphascon (Diphascon) serratum Pilato, Binda, Bertolani and Lisi, 2005
Species: Diphascon (Diphascon) sexbullatum Ito, 1995
Species: Diphascon (Diphascon) stappersi Richters, 1911
Species: Diphascon (Diphascon) tenue Thulin, 1928
Species: Diphascon (Diphascon) trachydorsatum Bartos, 1937
Species: Diphascon (Diphascon) victoriae Pilato and Binda, 1999
Species: Diphascon (Diphascon) zaniewi Kaczmarek and Michalczyk, 2004
Species: Diphascon bicorne Mihelcic, 1971
Species: Diphascon coniferens Bartos, 1960
Species: Diphascon marcuzzii Mihelcic, 1971
Species: Diphascon mariae Mihelcic, 1953
Species: Diphascon punctatum Iharos, 1963
Species: Diphascon rivulare Mihelcic, 1967
Species: Diphascon speciosum Mihelcic, 1971
Genus: Paradiphascon Dastych, 1992
Species: Paradiphascon manningi Dastych, 1992
Subfamilia: Hypsibiinae Pilato, 1969
Genus: Acutuncus Pilato & Binda, 1997
Species: Acutuncus antarcticus Richters, 1904
Synonym: Hypsibius simoizumii Sudzuki, 1964


Genus: Borealibius Pilato, Guidetti, Rebecchi, Lisi, Hansen & Bertolani, 2006
Species: Borealibius zetlandicus Murray, 1907
Genus: Hypsibius Ehrenberg, 1848
Species: Hypsibius allisoni Horning, Schuster & Grigarick, 1978


Species: Hypsibius antonovae Biserov, 1990
Species: Hypsibius arcticus Murray, 1907
Synonym: Macrobiotus heinisi Richters, 1907
Species: Hypsibius biscuitiformis Bartoš, 1960
Species: Hypsibius calcaratus Bartoš, 1935
Species: Hypsibius camelopardalis Ramazzotti & Maucci, 1983
Species: Hypsibius choucoutiensis Rahm, 1937
Species: Hypsibius conifer Mihelcic, 1938
Species: Hypsibius convergens Urbanowicz, 1925
Species: Hypsibius dujardini Doyère, 1840
Synonym: Macrobiotus palustris Dujardin, 1851
Synonym: Macrobiotus lacustris Dujardin, 1851
Synonym: Macrobiotus tetradactylus Lance, 1896
Synonym: Macrobiotus murrayi Richters, 1907
Synonym: Macrobiotus samoanus Richters, 1908
Synonym: Macrobiotus breckneri Richters, 1910
Synonym: Macrobiotus ursellus Della Valle, 1915
Synonym: Hypsibius murrayi Marcus, 1929
Species: Hypsibius fuhrmanni Heinis, 1914
Species: Hypsibius giusepperamazzotti Sudzuki, 1975
Species: Hypsibius heardensis Miller, McInnes & Bergstrom, 2005
Species: Hypsibius hypostomus Bartoš, 1935
Species: Hypsibius iskandarovi Tumanov, 1997
Species: Hypsibius janetscheki Ramazzotti, 1968
Species: Hypsibius klebelsbergi Mihelcic, 1959
Species: Hypsibius kunmingensis Yang, 2002
Species: Hypsibius macrocalcaratus Beasley, 1988
Species: Hypsibius maculatus Iharos, 1969
Species: Hypsibius marcelli Pilato, 1990
Species: Hypsibius microps Thulin, 1928 <Amended by Kaczmarek & Michalczyk 2009>
Species: Hypsibius montanus Iharos, 1940
Synonym: Hypsibius iharosi Bartoš, 1941
Species: Hypsibius morikawai Ito, 1995
Species: Hypsibius multituberculatus Pilato, Binda & Lisi, 2003
Species: Hypsibius novaezeelandiae Pilato & Binda, 1997
Species: Hypsibius pachyunguis Maucci, 1996
Species: Hypsibius pallidus Thulin, 1911 <Amended by Kaczmarek & Michalczyk 2009>
Species: Hypsibius pallidoides Pilato, Kiosya, Lisi, Inshina & Biserov, 2011
Species: Hypsibius pedrottii Bertolani, Manicardi & Gibertoni, 1987
Species: Hypsibius pradellii Bertolani & Rebecchi, 1996
Species: Hypsibius ragonesei Binda & Pilato, 1985
Species: Hypsibius roanensis Nelson & McGlothlin, 1993
Species: Hypsibius runae Bartoš, 1941
Species: Hypsibius scaber Maucci, 1987
Species: Hypsibius scabropygus Cuénot, 1929
Species: Hypsibius septulatus Pilato, Binda, Napolitano & Moncada, 2004
Species: Hypsibius seychellensis Pilato, Binda & Lisi, 2006
Species: Hypsibius shaanxiensis H. Li & X. Li, 2008
Species: Hypsibius stiliferus Abe, 2004
Species: Hypsibius thaleri Dastych, 2004
Subfamilia: Itaquasconinae Pilato, 1969
Genus: Astatumen Pilato, 1997
Species: Astatumen bartosi Weglarska, 1959
Species: Astatumen tamaensis Sudzuki, 1975
Species: Astatumen tamurai Ito, 1990
Species: Astatumen trinacriae Arcidiacono, 1962
Synonym: Itaquascon ramazzottii Iharos, 1966
Genus: Itaquascon De Barros, 1939
Species: Itaquascon biserovi Pilato, Binda & Moncada, 1999
Species: Itaquascon cambewarrense Pilato, Binda & Claxton, 2002
Species: Itaquascon enckelli Mihelcic, 1971/72


Species: Itaquascon globuliferum Abe & Ito, 1994
Species: Itaquascon mongolicus Kaczmarek, Michalczyk & Weglarska, 2002
Species: Itaquascon pawlowskii Weglarska, 1973
Species: Itaquascon pisoniae Pilato & Lisi, 2009
Species: Itaquascon placophorum Maucci, 1973
Species: Itaquascon simplex Mihelcic, 1971
Species: Itaquascon umbellinae de Barros, 1939
Species: Itaquascon unguiculum Pilato, Binda & Claxton, 2002
Genus: Mesocrista Pilato, 1987
Species: Mesocrista marcusi Rudescu, 1964
Species: Mesocrista spitzbergensis Richters, 1903



Genus: Parascon Pilato & Binda, 1987
Species: Parascon nichollsae Pilato & Lisi, 2004
Species: Parascon schusteri Pilato & Binda, 1987
Genus: Platicrista Pilato, 1987
Species: Platicrista affine Mihelcic, 1951
Species: Platicrista angustata Murray, 1905



Species: Platicrista cheleusis Kathman, 1990
Synonym: Diphascon craigi Beasley, 1990
Species: Platicrista horribilis Kaczmarek & Michalczyk, 2003
Species: Platicrista itaquasconoide Durante Pasa & Maucci, 1975
Species: Platicrista ramsayi Marley, 2006
Familia: Microhypsibiidae Pilato, 1998
Genus: Fractonotus Pilato, 1998
Species: Fractonotus caelatus Marcus, 1928



Genus: Microhypsibius Thulin, 1928
Species: Microhypsibius bertolanii Kristensen, 1982
Species: Microhypsibius japonicus Ito, 1991
Species: Microhypsibius minimus Kristensen, 1982
Species: Microhypsibius truncatus Thulin, 1928


Familia: Ramazzottidae Marley, McInnes & Sands, 2011
Genus: Hebesuncus Pilato, 1987
Species: Hebesuncus conjungens Thulin, 1911


Species: Hebesuncus ryani Dastych and Harris, 1994
Species: Hebesuncus schusteri Dastych, 1984
Genus: Ramazzottius Binda & Pilato, 1987
Species: Ramazzottius affinis Bertolani, Guidetti & Rebecchi, 1994
Species: Ramazzottius agannae Dastych, 2011
Species: Ramazzottius andreevi Biserov, 1997/98
Species: Ramazzottius anomalus Ramazzotti, 1962
Species: Ramazzottius baumanni Ramazzotti, 1962
Species: Ramazzottius belubellus Bartels, Nelson, Kaczmarek & Michalczyk, 2011
Species: Ramazzottius bunikowskae Kaczmarek, Michalczyk & Diduszko, 2006
Species: Ramazzottius cataphractus Maucci, 1974
Species: Ramazzottius caucasicus Biserov, 1997/98
Species: Ramazzottius edmondabouti Séméria, 1993
Species: Ramazzottius horningi Binda & Pilato, 1994
Species: Ramazzottius ljudmilae Biserov, 1997/98
Species: Ramazzottius montivagus Dastych, 1983
Species: Ramazzottius nivalis Dastych, 2006
Species: Ramazzottius novemcinctus Marcus, 1936
Species: Ramazzottius oberhaeuseri Doyère, 1840



Species: Ramazzottius rupeus Biserov, 1999
Species: Ramazzottius saltensis Claps & Rossi, 1984
Species: Ramazzottius semisculptus Pilato & Rebecchi, 1992
Species: Ramazzottius subanomalus Biserov, 1985
Species: Ramazzottius szeptycki Dastych, 1980 <Amended by Dastych 2009>


Species: Ramazzottius theroni Dastych, 1993
Species: Ramazzottius thulini Pilato, 1970
Species: Ramazzottius tribulosus Bertolani & Rebecchi, 1988
Species: Ramazzottius valaamis Biserov & Tumanov, 1993
Species: Ramazzottius varieornatus Bertolani & Kinchin, 1993
Superfamilia: Isohypsibioidea Marley, McInnes & Sands, 2011
Familia: Isohypsibiidae Marley, McInnes & Sands, 2011
Genus: Doryphoribius Pilato, 1969
Species: Doryphoribius amazzonicus Lisi, 2011
Species: Doryphoribius bertolanii Beasley and Pilato, 1987
Species: Doryphoribius bindae Lisi, 2011
Species: Doryphoribius dawkinsi Michalczyk & Kaczmarek, 2010
Species: Doryphoribius doryphorus Binda and Pilato, 1969
Species: Doryphoribius dupliglobulatus Ito, 1995
Species: Doryphoribius evelinae Marcus, 1928
Species: Doryphoribius flavus Iharos, 1966
Synonym: Doryphoribius citrinus Maucci, 1973
Species: Doryphoribius gibber Beasley and Pilato, 1987
Species: Doryphoribius huangguoshuensis H. Wang, L. Wang & X. Li, 2007
Species: Doryphoribius koreanus Moon, Kim and Bertolani, 1994
Species: Doryphoribius korganovae Biserov, 1994
Species: Doryphoribius longistipes Bartels, Nelson, Kaczmarek & Michalczyk, 2008
Species: Doryphoribius macrodon Binda, Pilato and Dastych, 1980
Species: Doryphoribius maranguensis Binda and Pilato, 1995
Species: Doryphoribius mariae Pilato and Binda, 1991
Species: Doryphoribius mexicanus Beasley, Kaczmarek & Michalczyk, 2008
Species: Doryphoribius minimus Bartels, Nelson, Kaczmarek & Michalczyk, 2008
Species: Doryphoribius neglectus Pilato and Lisi, 2004
Species: Doryphoribius picoensis Fontoura, Pilato & Lisi, 2008
Species: Doryphoribius pilatoi Bertolani, 1984
Species: Doryphoribius polynettae Biserov, 1988
Species: Doryphoribius qinlingense X. Li, Su and Yu, 2004
Species: Doryphoribius quadrituberculatus Kaczmarek and Michalczyk, 2004
Species: Doryphoribius smokiensis Bartels, Nelson, Kaczmarek & Michalczyk, 2007
Species: Doryphoribius solidunguis Lisi, 2011
Species: Doryphoribius taiwanus X. Li & H. Li, 2008
Species: Doryphoribius tergumrudis Bartels, Nelson, Kaczmarek & Michalczyk, 2008
Species: Doryphoribius tessellatus Meyer, 2011
Species: Doryphoribius turkmenicus Biserov, 1999
Species: Doryphoribius vietnamensis Iharos, 1969
Species: Doryphoribius zappalai Pilato, 1971
Species: Doryphoribius zyxiglobus Horning, Schuster and Grigarick, 1978
Genus: Eremobiotus Biserov, 1992
Species: Eremobiotus alicatai Binda, 1969
Species: Eremobiotus ovezovae Biserov, 1992
Genus: Halobiotus Kristensen, 1982
Species: Halobiotus arcturulius Kristensen, 1983
Species: Halobiotus crispae Kristensen, 1982
Species: Halobiotus stenostomus Richters, 1908
Synonym: Macrobiotus appelloefi Richters, 1908
Synonym: Hypsibius (Isohypsibius) geddesi Hallas, 1971




Genus: Isohypsibius Thulin, 1928
Species: Isohypsibius altai Kaczmarek & Michalczyk, 2006
Species: Isohypsibius annulatus Murray, 1905
Subspecies: Isohypsibius annulatus annulatus Murray, 1905
Subspecies: Isohypsibius annulatus minor Ramazzotti, 1945
Species: Isohypsibius arbiter Binda, 1980
Species: Isohypsibius archangajensis Kaczmarek & Michalczyk, 2004
Species: Isohypsibius arcuatus Bartoš, 1934
Species: Isohypsibius asper Murray, 1906
Species: Isohypsibius austriacus Iharos, 1966
Species: Isohypsibius baicalensis Ramazzotti, 1966
Species: Isohypsibius baldii Ramazzotti, 1945
Species: Isohypsibius baldiioides Tumanov, 2003
Species: Isohypsibius barbarae Pilato & Binda, 2002
Species: Isohypsibius bartosi Iharos, 1966
Species: Isohypsibius basalovoi Durante & Maucci, 1973
Species: Isohypsibius belliformis Mihelcic, 1971
Species: Isohypsibius bellus Mihelcic, 1971
Species: Isohypsibius borkini Tumanov, 2003
Species: Isohypsibius brevispinosus Iharos, 1966
Species: Isohypsibius brevitubulatus Rho, Chang & Kim, 1997
Species: Isohypsibius brulloi Pilato & Pennisi, 1976
Species: Isohypsibius bulbifer Mihelcic, 1957
Species: Isohypsibius cameruni Iharos, 1969
Species: Isohypsibius campbellensis Pilato, 1996
Species: Isohypsibius canadensis Murray, 1910


Species: Isohypsibius ceciliae Pilato & Binda, 1987
Species: Isohypsibius changbaiensis Yang, 1999
Species: Isohypsibius chiarae Maucci, 1987
Species: Isohypsibius costatus Mihelcic, 1971
Species: Isohypsibius cyrilli Mihelcic, 1951
Species: Isohypsibius damxungensis Yang, 2007
Species: Isohypsibius dastychi Pilato, Bertolani & Binda, 1982
Species: Isohypsibius deconincki Pilato, 1971
Species: Isohypsibius deflexus Mihelcic, 1960
Species: Isohypsibius dudichi Iharos, 1964
Species: Isohypsibius duranteae Maucci, 1978
Species: Isohypsibius effusus Mihelcic, 1971
Species: Isohypsibius elegans Binda & Pilato, 1971
Species: Isohypsibius eplenyiensis Iharos, 1970
Species: Isohypsibius franzi Mihelcic, 1951
Species: Isohypsibius fuscus Mihelcic, 1971/72
Species: Isohypsibius gilvus Biserov, 1986
Species: Isohypsibius glaber Durante Pasa & Maucci, 1979
Species: Isohypsibius glazovi Biserov, 1999
Species: Isohypsibius gracilis Iharos, 1966
Species: Isohypsibius granditintinus Chang & Rho, 1996
Species: Isohypsibius granulifer Thulin, 1928
Subspecies: Isohypsibius granulifer granulifer Thulin, 1928
Subspecies: Isohypsibius granulifer koreanensis Iharos, 1971
Species: Isohypsibius gyulai Mihelcic, 1971
Species: Isohypsibius hadzii Mihelcic, 1938
Species: Isohypsibius helenae Iharos, 1964
Species: Isohypsibius hydrogogianus Ito & Tagami, 1993
Species: Isohypsibius hypostomoides Mihelcic, 1971
Species: Isohypsibius improvisus Dastych, 1984
Species: Isohypsibius indicus Murray, 1907
Species: Isohypsibius irregibilis Biserov, 1992
Species: Isohypsibius jakieli Dastych, 1984
Species: Isohypsibius jingshanensis Yang, 2003
Species: Isohypsibius jinhouensis Yang, 2007
Species: Isohypsibius josephi Iharos, 1964
Species: Isohypsibius kenodontis Kendall-Fite & Nelson, 1996
Species: Isohypsibius kotovae Tumanov, 2003
Species: Isohypsibius kristenseni Pilato, Catanzaro & Binda, 1989
Species: Isohypsibius ladogensis Tumanov, 2003
Species: Isohypsibius laevis McInnes, 1995
Species: Isohypsibius latiunguis Iharos, 1964
Species: Isohypsibius leithaicus Iharos, 1966
Species: Isohypsibius liae X. Li & L. Wang, 2006
Species: Isohypsibius lineatus Mihelcic, 1969
Species: Isohypsibius longiunguis Pilato, 1974
Species: Isohypsibius lunulatus Iharos, 1966
Species: Isohypsibius macrodactylus Maucci, 1978
Synonym: Isohypsibius zierhofferi Dastych, 1980
Species: Isohypsibius malawiensis Jørgensen, 2001
Species: Isohypsibius mammillosus Iharos, 1964
Species: Isohypsibius marcellinoi Binda & Pilato, 1971
Species: Isohypsibius marii Bertolani, 1982
Species: Isohypsibius mihelcici Iharos, 1964
Species: Isohypsibius monoicus Bertolani, 1982
Species: Isohypsibius monstruosus Maucci, 1991
Species: Isohypsibius montanus Mihelcic, 1938
Species: Isohypsibius myrops du Bois-Reymond Marcus, 1944
Species: Isohypsibius neoundulatus Durante Pasa & Maucci, 1975
Species: Isohypsibius nipponicus Sudzuki, 1975
Species: Isohypsibius nodosus Murray, 1907
Species: Isohypsibius novaeguineae Iharos, 1967
Species: Isohypsibius palmai Pilato, 1996
Species: Isohypsibius panovi Tumanov, 2005
Species: Isohypsibius papillifer Murray, 1905
Subspecies: Isohypsibius papillifer bulbosus Marcus, 1928
Subspecies: Isohypsibius papillifer indicus Iharos, 1969
Subspecies: Isohypsibius papillifer papillifer Murray, 1905
Species: Isohypsibius pappi Iharos, 1966
Species: Isohypsibius pauper Mihelcic, 1971
Species: Isohypsibius pilatoi Durante Pasa & Maucci, 1979
Species: Isohypsibius pratensis Iharos, 1964
Species: Isohypsibius prosostomus Thulin, 1928
Subspecies: Isohypsibius prosostomus cambrensis Morgan, 1976
Subspecies: Isohypsibius prosostomus prosostomus Thulin, 1928
Species: Isohypsibius pseudoundulatus da Cunha & do Nascimento Ribeiro, 1964
Species: Isohypsibius pulcher Mihelcic, 1971/72
Species: Isohypsibius pushkini Tumanov, 2003
Species: Isohypsibius qinlingensis X. Li, L. Wang & Yu, 2005
Species: Isohypsibius rahmi X. Li & L. Wang, 2006
Species: Isohypsibius reticulatus Pilato, 1973
Species: Isohypsibius roberti Biserov, 1996
Species: Isohypsibius ronsisvallei Binda & Pilato, 1969
Species: Isohypsibius rudescui Iharos, 1966
Species: Isohypsibius rugosus Guidi & Grabowski, 1996
Species: Isohypsibius sabellai Pilato, Binda, Napolitano & Moncada, 2004
Species: Isohypsibius sattleri Richters, 1902
Synonym: Hypsibius bakonyiensis Iharos, 1964
Synonym: Isohypsibius bakonyiensis Iharos, 1964
Species: Isohypsibius schaudinni Richters, 1909
Species: Isohypsibius sculptus Ramazzotti, 1962
Species: Isohypsibius sellnicki Mihelcic, 1962
Species: Isohypsibius septentrionalis Thulin, 1928
Species: Isohypsibius silvicola Iharos, 1966
Species: Isohypsibius sismicus Maucci, 1978
Species: Isohypsibius solidus Mihelcic, 1971
Species: Isohypsibius taibaiensis X. Li & L. Wang, 2005
Species: Isohypsibius tetradactyloides Richters, 1907
Species: Isohypsibius theresiae Iharos, 1964
Species: Isohypsibius torulosus Mihelcic, 1959
Species: Isohypsibius truncorum Iharos, 1964
Species: Isohypsibius tuberculatus Plate, 1888
Species: Isohypsibius tuberculoides Mihelcic, 1951
Species: Isohypsibius tubereticulatus Pilato & Catanzaro, 1989
Species: Isohypsibius tucumanensis Claps & Rossi, 1984
Species: Isohypsibius undulatus Thulin, 1928
Species: Isohypsibius vejdovskyi Bartoš, 1939
Species: Isohypsibius verae Pilato & Catanzaro, 1989
Species: Isohypsibius verrucosus Della Valle, 1915
Synonym: Isohypsibius gibbus Marcus, 1928
Species: Isohypsibius wilsoni Horning, Schuster & Grigarick, 1978
Species: Isohypsibius woodsae Kathman, 1990
Species: Isohypsibius yunnanensis Yang, 2002
Genus: Mixibius Pilato, 1992
Species: Mixibius fueginus Pilato & Binda, 1996
Species: Mixibius ninguidus Biserov, 1999
Species: Mixibius ornatus Pilato, Binda, Napolitano & Moncada, 2002
Species: Mixibius pilatoi L. Wang, 2009
Species: Mixibius saracenus Pilato, 1973
Species: Mixibius schnurae Pilato, Lisi & Binda, 2010
Species: Mixibius sutirae Pilato, Binda & Lisi, 2004
Species: Mixibius tibetanus H. Li & X. Li, 2008
Genus: Pseudobiotus Nelson, 1980
Species: Pseudobiotus hirsutellus Pilato, Lisi & Binda, 2010
Species: Pseudobiotus kathmanae Nelson, Marley & Bertolani, 1999
Species: Pseudobiotus longiunguis Iharos, 1968
Species: Pseudobiotus matici Pilato, 1971
Species: Pseudobiotus megalonyx Thulin, 1928
Species: Pseudobiotus spinifer Chang, Kaczmarek, Lee & Michalczyk, 2007
Species: Pseudobiotus vladimiri Biserov, Dudichev & Biserova, 2001 <Amended by Chang et al. 2007>
Genus: Ramajendas Pilato & Binda, 1991
Species: Ramajendas frigidus Pilato & Binda, 1991
Species: Ramajendas heatwolei Miller, Horning & Dastych, 1995
Species: Ramajendas renaudi Ramazzotti, 1972


Genus: Thalerius Dastych, 2009
Species: Thalerius konradi Dastych, 2009
Genus: Thulinius Bertolani, 2003
Synonym: Thulinia Bertolani, 1981
Species: Thulinius augusti Murray, 1907



Synonym: Macrobiotus lacustris Wenck, 1914
Species: Thulinius itoi Tsurusaki, 1980
Species: Thulinius ruffoi Bertolani, 1982
Species: Thulinius saltursus Schuster, Toftner & Grigarick, 1978
Species: Thulinius stephaniae Pilato, 1974


Superfamilia: Macrobiotoidea Thulin, 1928 in Marley et al. 2011
Familia: Macrobiotidae Thulin, 1928
Genus: Adorybiotus Maucci & Ramazzotti, 1981
Species: Adorybiotus granulatus Richters, 1903
Genus: Biserovus Guidetti & Pilato, 2003
Species: Biserovus bindae Christenberry & Higgins, 1979
Genus: Calcarobiotus Dastych, 1993
Subgenus: Calcarobiotus (Calcarobiotus) Dastych, 1993
Species: Calcarobiotus (Calcarobiotus) digeronimoi Pilato, Binda & Lisi, 2004
Species: Calcarobiotus (Calcarobiotus) filmeri Dastych, 1993
Species: Calcarobiotus (Calcarobiotus) gildae Maucci & Durante Pasa, 1980
Species: Calcarobiotus (Calcarobiotus) hainanensis X. Li, D. Wang & L. Wang, 2008
Species: Calcarobiotus (Calcarobiotus) imperialis Abe & Takeda, 2000
Species: Calcarobiotus (Calcarobiotus) longinoi Kaczmarek, Michalczyk & Guidetti, 2006
Species: Calcarobiotus (Calcarobiotus) occultus Dastych, 1993
Species: Calcarobiotus (Calcarobiotus) parvicalcar Pilato & Lisi, 2009
Subgenus: Calcarobiotus (Discrepunguis) Guidetti & Bertolani, 2001
Species: Calcarobiotus (Discrepunguis) polygonatus Binda & Guglielmino, 1991
Synonym: Macrobiotus eugranulatus Maucci, 1993
Species: Calcarobiotus (Discrepunguis) tetrannulatus Pilato, Binda & Lisi, 2004
Genus: Famelobiotus Pilato, Binda & Lisi, 2004
Species: Famelobiotus scalicii Pilato, Binda & Lisi, 2004
Genus: Insuetifurca Guidetti & Pilato, 2003
Species: Insuetifurca arrowsmithi Kathman & Nelson, 1989
Species: Insuetifurca austronipponica Abe, 2005
Species: Insuetifurca fujiense Ito, 1997
Species: Insuetifurca xiae X. Li, Su & Yu, 2004 <Amended and transferred from Biserovus to Insuetifurca by Li 2009>
Genus: Macrobiotus C.A.S. Schultze, 1834
Species: Macrobiotus almadai Fontoura, Pilato & Lisi, 2008
Species: Macrobiotus altitudinalis Biserov, 1997/98
Species: Macrobiotus alvaroi Pilato & Kaczmarek, 2007
Species: Macrobiotus anderssoni Richters, 1907
Species: Macrobiotus andinus Maucci, 1988
Species: Macrobiotus annae Richters, 1908
Species: Macrobiotus aradasi Binda, Pilato & Lisi, 2005
Species: Macrobiotus arguei Pilato & Sperlinga, 1975
Species: Macrobiotus ariekammensis Weglarska, 1965
Synonym: Macrobiotus adelges Dastych, 1977
Species: Macrobiotus armatus Pilato & Binda, 1996
Species: Macrobiotus artipharyngis Iharos, 1940
Species: Macrobiotus ascensionis Richters, 1908
Species: Macrobiotus australis Pilato & D'Urso, 1976
Species: Macrobiotus baltatus McInnes, 1991
Species: Macrobiotus barabanovi Tumanov, 2005
Species: Macrobiotus binieki Kaczmarek, Gołdyn, Prokop & Michalczyk, 2011
Species: Macrobiotus barbarae Kaczmarek, Michalczyk & Degma, 2007
Species: Macrobiotus biserovi Bertolani, Guidi & Rebecchi, 1996
Species: Macrobiotus blocki Dastych, 1984
Species: Macrobiotus brevipes Mihelcic, 1971/72
Species: Macrobiotus caelicola Kathman, 1990
Species: Macrobiotus carsicus Maucci, 1954
Species: Macrobiotus caymanensis Meyer, 2011
Species: Macrobiotus contii Pilato & Lisi, 2006
Species: Macrobiotus coronatus de Barros, 1942
Species: Macrobiotus creber Pilato & Lisi, 2009
Species: Macrobiotus crenulatus Richters, 1904
Synonym: Macrobiotus dentatus Binda, 1974
Species: Macrobiotus danielisae Pilato, Binda & Lisi, 2006
Species: Macrobiotus dariae Pilato & Bertolani, 2004
Species: Macrobiotus denticulus Dastych, 2002
Species: Macrobiotus diffusus Binda & Pilato, 1987
Species: Macrobiotus diguensis Pilato & Lisi, 2009
Species: Macrobiotus dimentmani Pilato, Lisi & Binda, 2010
Species: Macrobiotus divergens Binda, Pilato & Lisi, 2005
Species: Macrobiotus diversus Biserov, 1990
Species: Macrobiotus drakensbergi Dastych, 1993
Species: Macrobiotus echinogenitus Richters, 1904
Species: Macrobiotus erminiae Binda & Pilato, 1999
Species: Macrobiotus evelinae de Barros, 1938
Species: Macrobiotus furciger Murray, 1907
Synonym: Macrobiotus ehrenbergi Heinis, 1921
Synonym: Macrobiotus furcatus Murray, 1906
Species: Macrobiotus gemmatus Bartoš, 1963
Species: Macrobiotus glebkai Biserov, 1990
Species: Macrobiotus grandis Richters, 1911
Species: Macrobiotus halei Bartels, Pilato, Lisi & Nelson, 2009
Species: Macrobiotus hapukuensis Pilato, Binda & Lisi, 2006
Species: Macrobiotus harmsworthi harmsworthi Murray, 1907
Subspecies: Macrobiotus harmsworthi harmsworthi Murray, 1907
Subspecies: Macrobiotus harmsworthi obscurus Dastych, 1985
Species: Macrobiotus hibiscus de Barros, 1942
Species: Macrobiotus hieronimi Pilato & Claxton, 1988
Species: Macrobiotus hufelandi C.A.S. Schultze, 1833
Subspecies: Macrobiotus hufelandi hufelandi C.A.S. Schultze, 1833
Subspecies: Macrobiotus hufelandi maculatus Iharos, 1973
Species: Macrobiotus humilis Binda & Pilato, 2001
Species: Macrobiotus hyperboreus Biserov, 1990
Species: Macrobiotus iharosi Pilato, Binda & Catanzaro, 1991
Species: Macrobiotus insignis Bartoš, 1963
Species: Macrobiotus insularis Pilato, 2006
Species: Macrobiotus islandicus Richters, 1904
Subspecies: Macrobiotus islandicus islandicus Richters, 1904
Synonym: Macrobiotus ruffoi Maucci, 1973
Subspecies: Macrobiotus islandicus nicaraguensis Séméria, 1985
Species: Macrobiotus joannae Pilato & Binda, 1983
Species: Macrobiotus kazmierskii Kaczmarek & Michalczyk, 2009
Species: Macrobiotus kirghizicus Tumanov, 2005
Species: Macrobiotus kolleri Mihelcic, 1951
Species: Macrobiotus komareki Bartoš, 1939
Species: Macrobiotus kovalevi Tumanov, 2004
Species: Macrobiotus krynauwi Dastych & Harris, 1995
Species: Macrobiotus kurasi Dastych, 1981
Species: Macrobiotus lazzaroi Maucci, 1986
Synonym: Macrobiotus spallanzanii Maucci, 1973
Species: Macrobiotus lissostomus Durante Pasa & Maucci, 1979
Species: Macrobiotus liviae Ramazzotti, 1962
Species: Macrobiotus longipes Mihelcic, 1971/72
Species: Macrobiotus lusitanicus Maucci & Durante Pasa, 1984
Species: Macrobiotus macrocalix Bertolani & Rebecchi, 1993
Species: Macrobiotus madegassus Maucci, 1993
Species: Macrobiotus mandalaae Pilato, 1974
Species: Macrobiotus marlenae Kaczmarek & Michalczyk, 2004
Species: Macrobiotus martini Bartels, Pilato, Lisi & Nelson, 2009
Species: Macrobiotus mauccii Pilato, 1974
Species: Macrobiotus meridionalis Richters, 1909
Species: Macrobiotus modestus Pilato & Lisi, 2009
Species: Macrobiotus montanus Murray, 1910
Synonym: Macrobiotus morulatus Bartoš, 1936
Species: Macrobiotus mottai Binda & Pilato, 1994
Species: Macrobiotus nelsonae Guidetti, 1998
Species: Macrobiotus neuquensis Rossi, Claps & Ardohain, 2009
Species: Macrobiotus norvegicus Mihelcic, 1971/72
Species: Macrobiotus nuragicus Pilato & Sperlinga, 1975
Species: Macrobiotus occidentalis Murray, 1910
Subspecies: Macrobiotus occidentalis occidentalis Murray, 1910
Subspecies: Macrobiotus occidentalis striatus Dastych, 1974
Species: Macrobiotus ocotensis Pilato, 2006
Species: Macrobiotus orcadensis Murray, 1907
Species: Macrobiotus ovidii Bartoš, 1937
Species: Macrobiotus ovostriatus Pilato & Patanè, 1998
Species: Macrobiotus ovovillosus Baumann, 1960
Species: Macrobiotus pallarii Maucci, 1954
Synonym: Macrobiotus aviglianae Robotti, 1970
Species: Macrobiotus papillosus Iharos, 1963
Species: Macrobiotus patagonicus Maucci, 1988
Species: Macrobiotus patiens Pilato, Binda, Napolitano & Moncada, 2000
Species: Macrobiotus perfidus Pilato & Lisi, 2009
Species: Macrobiotus persimilis Binda & Pilato, 1972
Species: Macrobiotus personatus Biserov, 1990
Species: Macrobiotus peterseni Maucci, 1991
Species: Macrobiotus pilatoi Binda & Rebecchi, 1992
Species: Macrobiotus polaris Murray, 1910
Species: Macrobiotus polonicus Pilato, Kaczmarek, Michalczyk & Lisi, 2003
Species: Macrobiotus polyopus Marcus, 1928
Species: Macrobiotus porteri Rahm, 1931
Species: Macrobiotus potockii Weglarska, 1968
Species: Macrobiotus primitivae de Barros, 1942
Species: Macrobiotus priviterae Binda, Pilato, Moncada & Napolitano, 2001
Species: Macrobiotus psephus du Bois-Reymond Marcus, 1944
Species: Macrobiotus pseudocoronatus Pilato, Binda & Lisi, 2006
Species: Macrobiotus pseudofurcatus Pilato, 1972
Species: Macrobiotus pseudoliviae Pilato & Binda, 1996
Species: Macrobiotus pseudonuragicus Pilato, Binda & Lisi, 2004
Species: Macrobiotus punctillus Pilato, Binda & Azzaro, 1990
Species: Macrobiotus radiatus Pilato, Binda & Catanzaro, 1991
Species: Macrobiotus ragonesei Binda, Pilato, Moncada & Napolitano, 2001
Species: Macrobiotus ramoli Dastych, 2005
Species: Macrobiotus rawsoni Horning, Schuster & Grigarick, 1978
Species: Macrobiotus recens Cuénot, 1932
Species: Macrobiotus reinhardti Michalczyk & Kaczmarek, 2003
Species: Macrobiotus rigidus Pilato & Lisi, 2006
Species: Macrobiotus rollei Heinis, 1921
Species: Macrobiotus rubens Murray, 1907
Species: Macrobiotus sandrae Bertolani & Rebecchi, 1993
Species: Macrobiotus santoroi Pilato & D'Urso, 1976
Species: Macrobiotus sapiens Binda & Pilato, 1984
Species: Macrobiotus semmelweisi Pilato, Binda & Lisi, 2006
Species: Macrobiotus serratus Bertolani, Guidi & Rebecchi, 1996
Species: Macrobiotus seychellensis Biserov, 1994
Species: Macrobiotus shennongensis Yang, 1999
Species: Macrobiotus siamensis Tumanov, 2006
Species: Macrobiotus sicheli Binda, Pilato & Lisi, 2005
Species: Macrobiotus simulans Pilato, Binda, Napolitano & Moncada, 2000
Species: Macrobiotus sklodowskae Michalczyk, Kaczmarek & Weglarska, 2006
Species: Macrobiotus snaresensis Horning, Schuster & Grigarick, 1978
Species: Macrobiotus spectabilis Thulin, 1928
Species: Macrobiotus spertii Ramazzotti, 1957
Species: Macrobiotus stellaris du Bois-Reymond Marcus, 1944
Species: Macrobiotus striatus Mihelcic, 1949
Species: Macrobiotus submorulatus Iharos, 1966
Species: Macrobiotus szeptyckii Kaczmarek & Michalczyk, 2009
Species: Macrobiotus tehuelchensis Rossi, Claps & Ardohain, 2009
Species: Macrobiotus terminalis Bertolani & Rebecchi, 1993
Species: Macrobiotus terricola Mihelcic, 1951
Species: Macrobiotus tetraplacoides Fontoura, 1981
Species: Macrobiotus topali Iharos, 1969
Species: Macrobiotus trunovae Biserov, Pilato & Lisi, 2011
Species: Macrobiotus virgatus Murray, 1910
Species: Macrobiotus vladimiri Bertolani, Biserov, Rebecchi & Cesari, 2011
Species: Macrobiotus wauensis Iharos, 1973
Species: Macrobiotus wuzhishanensis Yin, L. Wang & X. Li, 2011
Species: Macrobiotus yunshanensis Yang, 2002
Species: Macrobiotus zhejiangensis Yin, L. Wang & X. Li, 2011
Genus: Minibiotus R.O. Schuster, 1980
Species: Minibiotus acadianus Meyer & Domingue, 2011
Species: Minibiotus acontistus de Barros, 1942
Species: Minibiotus aculeatus Murray, 1910
Synonym: Macrobiotus intermedius subjulietae Horning, Schuster & Grigarick, 1978
Species: Minibiotus africanus Binda & Pilato, 1995
Species: Minibiotus allani Murray, 1913
Species: Minibiotus aquatilis Claxton, 1998
Species: Minibiotus asteris Claxton, 1998
Species: Minibiotus bisoctus Horning, Schuster & Grigarick, 1978
Species: Minibiotus claxtonae Rossi, Claps & Ardohain, 2009
Species: Minibiotus constellatus Michalczyk & Kaczmarek, 2003
Species: Minibiotus continuus Pilato & Lisi, 2006
Species: Minibiotus crassidens Murray, 1907
Species: Minibiotus decrescens Binda & Pilato, 1995
Species: Minibiotus diphasconides Iharos, 1969
Species: Minibiotus eichhorni Michalczyk & Kaczmarek, 2004
Species: Minibiotus ethelae Claxton, 1998
Species: Minibiotus fallax Pilato, Claxton & Binda, 1989
Species: Minibiotus floriparus Claxton, 1998
Species: Minibiotus furcatus Ehrenberg, 1859
Synonym: Macrobiotus furcatus Murray, 1906
Species: Minibiotus granatai Pardi, 1941
Species: Minibiotus gumersindoi Guil & Guidetti, 2005
Species: Minibiotus harrylewisi Meyer & Hinton, 2009
Species: Minibiotus hispidus Claxton, 1998
Species: Minibiotus hufelandioides Murray, 1910
Species: Minibiotus intermedius Plate, 1888
Species: Minibiotus jonesorum Meyer, Lyons, Nelson & Hinton, 2011
Species: Minibiotus julietae de Barros, 1942
Species: Minibiotus keppelensis Claxton, 1998
Species: Minibiotus maculartus Pilato & Claxton, 1988
Species: Minibiotus marcusi de Barros, 1942
Species: Minibiotus milleri Claxton, 1998
Species: Minibiotus orthofasciatus Fontoura, Pilato, Lisi & Morais, 2009
Species: Minibiotus pilatus Claxton, 1998
Species: Minibiotus poricinctus Claxton, 1998
Species: Minibiotus pustulatus Ramazzotti, 1959
Species: Minibiotus ramazzottii Binda & Pilato, 1992
Species: Minibiotus scopulus Claxton, 1998
Species: Minibiotus sidereus Pilato, Binda & Lisi, 2003
Species: Minibiotus stuckenbergi Dastych, Ryan & Watkins, 1990
Species: Minibiotus subintermedius Ramazzotti, 1962
Species: Minibiotus taiti Claxton, 1998
Species: Minibiotus vinciguerrae Binda & Pilato, 1992
Species: Minibiotus weglarskae Michalczyk, Kaczmarek & Claxton, 2005
Species: Minibiotus weinerorum Dastych, 1984
Species: Minibiotus wuzhishanensis X. Li, D. Wang & L. Wang, 2008
Species: Minibiotus xavieri Fontoura, Pilato, Morais & Lisi, 2009
Genus: Minilentus Guidetti & Pilato, 2003
Species: Minilentus dubius Schuster & Toftner, 1982
Genus: Paramacrobiotus Guidetti, Schill, Bertolani, Dandekar & Wolf, 2009
Species: Paramacrobiotus alekseevi Tumanov, 2005 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus areolatus Murray, 1907 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus beotiae Durante Pasa & Maucci, 1979 {species dubia; Guidetti et al. 2009}
Species: Paramacrobiotus centesimus Pilato, 2000 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus chieregoi Maucci & Durante Pasa, 1980 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus corgatensis Pilato, Binda & Lisi, 2002 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus crenatus Maucci, 1991 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus csotiensis Iharos, 1966 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus danielae Pilato, Binda, Napolitano & Moncada, 2001 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus derkai Degma, Michalczyk & Kaczmarek, 2008 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus fairbanksi Schill, Förster, Dandekar & Wolf, 2010
Species: Paramacrobiotus garynahi Kaczmarek, Michalczyk & Diduszko, 2005 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus gerlachae Pilato, Binda & Lisi, 2004 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus huziori Mihalczyk & Kaczmarek, 2006 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus kenianus Schill, Förster, Dandekar & Wolf, 2010
Species: Paramacrobiotus lorenae Biserov, 1996 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus magdalenae Mihalczyk & Kaczmarek, 2006 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus palaui Schill, Förster, Dandekar & Wolf, 2010
Species: Paramacrobiotus peteri Pilato, Claxton & Binda, 1989 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus richtersi Murray, 1911 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Synonym: Macrobiotus schultzei Greeff, 1866
Species: Paramacrobiotus rioplatensis Claps & Rossi, 1997 {doubly attributed to the genus; Guidetti et al. 2009}
Species: Paramacrobiotus savai Binda and Pilato, 2001 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus tonollii Ramazzotti, 1956 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus vanescens Pilato, Binda and Catanzaro, 1991 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Species: Paramacrobiotus walteri Biserov, 1997/98 <Transferred from Macrobiotus to Paramacrobiotus by Guidetti et al. 2009>
Genus: Pseudodiphascon Ramazzotti, 1965
Species: Pseudodiphascon inflexum Arcidiacono, 1964


Genus: Pseudohexapodibius Bertolani & Biserov, 1996
Species: Pseudohexapodibius degenerans Biserov, 1990
Genus: Richtersius Pilato & Binda, 1989
Synonym: Richtersia Pilato & Binda, 1987
Species: Richtersius coronifer Richters, 1903



Genus: Schusterius Kaczmarek & Michalczyk, 2006
Species: Schusterius tridigitus R.O. Schuster, 1983
Genus: Tenuibiotus Pilato & Lisi, 2011
Species: Tenuibiotus bondavallii Manicardi, 1989
Species: Tenuibiotus bozhkae Pilato, Kiosya, Lisi, Inshina & Biserov, 2011
Species: Tenuibiotus ciprianoi Guil, Guidetti & Machordom, 2007
Species: Tenuibiotus danilovi Tumanov, 2007
Species: Tenuibiotus higginsi Maucci, 1987
Species: Tenuibiotus hyperonyx Maucci, 1983
Species: Tenuibiotus hystricogenitus Maucci, 1978
Species: Tenuibiotus kozharai Biserov, 1999
Species: Tenuibiotus mongolicus Maucci, 1988
Species: Tenuibiotus tenuiformis Tumanov, 2007
Species: Tenuibiotus tenuis Binda & Pilato, 1972
Species: Tenuibiotus voronkovi Tumanov, 2007
Species: Tenuibiotus willardi Pilato, 1977
Genus: Xerobiotus Bertolani & Biserov, 1996
Species: Xerobiotus euxinus Pilato, Kiosya, Lisi, Inshina & Biserov, 2011
Species: Xerobiotus pseudohufelandi Iharos, 1966
Synonym: Macrobiotus inermis Binda & Pilato, 1971
Species: Xerobiotus xerophilus Dastych, 1978
Familia: Murrayidae Guidetti, Gandolfi, Rossi & Bertolani, 2005
Genus: Dactylobiotus Schuster, 1980
Species: Dactylobiotus ambiguus Murray, 1907
Species: Dactylobiotus ampullaceus Thulin, 1911
Species: Dactylobiotus aquatilis Yang, 1999
Species: Dactylobiotus caldarellai Pilato & Binda, 1994
Species: Dactylobiotus dervizi Biserov, 1998
Species: Dactylobiotus dispar Murray, 1907
Species: Dactylobiotus grandipes Schuster, Toftner & Grigarick, 1978
Species: Dactylobiotus haplonyx Maucci, 1981
Species: Dactylobiotus henanensis Yang, 2002
Species: Dactylobiotus kansae Beasley, Miller & Shively, 2009
Species: Dactylobiotus lombardoi Binda & Pilato, 1999
Species: Dactylobiotus luci Kaczmarek, Michalczyk & Eggermont, 2008
Species: Dactylobiotus macronyx Dujardin, 1851
Species: Dactylobiotus octavi Guidetti, Altiero & Hansen, 2006
Species: Dactylobiotus parthenogeneticus Bertolani, 1982
Species: Dactylobiotus selenicus Bertolani, 1982
Genus: Macroversum Pilato & Catanzaro, 1988
Species: Macroversum mirum Pilato & Catanzaro, 1988
Genus: Murrayon Bertolani & Pilato, 1988
Species: Murrayon dianeae Kristensen, 1982


Species: Murrayon hastatus Murray, 1907
Species: Murrayon hibernicus Murray, 1911
Species: Murrayon nocentiniae Ramazzotti, 1961
Species: Murrayon ovoglabellus Biserov, 1988
Species: Murrayon pullari Murray, 1907


Species: Murrayon stellatus Guidetti, 1998
Familia: Necopinatidae Ramazzotti and Maucci, 1983
Genus: Necopinatum Pilato, 1971
Species: Necopinatum mirabile Pilato, 1971
Familia: †Beornidae Cooper, 1964
Genus: Beorn Cooper, 1964
Species: Beorn leggi Cooper, 1964
Genus: Apodibius Dastych, 1983
Species: Apodibius confusus Dastych, 1983
Species: Apodibius nuntius Binda, 1984
Synonym: Apodibius serventyi Morgan & Nicholls, 1986
Species: Apodibius richardi Vargha, 1996
Classis: Heterotardigrada Marcus, 1927
Ordo: Arthrotardigrada Marcus, 1927
Familia: Archechiniscidae Binda, 1978
Synonym: Archechiniscinae De Zio Grimaldi & D'Addabbo Gallo, 1987
Genus: Archechiniscus Schulz, 1953
Species: Archechiniscus marci Schulz, 1953
Species: Archechiniscus minutus De Zio Grimaldi & D'Addabbo Gallo, 1987
Species: Archechiniscus symbalanus Chang & Rho, 1998
Familia: Batillipedidae Ramazzotti, 1963
Synonym: Discopodidae Marcus, 1934
Genus: Batillipes Richters, 1909
Species: Batillipes acaudatus Pollock, 1971
Species: Batillipes adriaticus De Zio Grimaldi, Morone De Lucia and D'Addabbo Gallo, 1979
Species: Batillipes africanus Morone De Lucia, D'Addabbo Gallo and De Zio Grimaldi, 1988
Species: Batillipes annulatus de Zio, 1962
Species: Batillipes bullacaudatus McGinty & Higgins, 1968
Species: Batillipes carnonensis Fize, 1957
Species: Batillipes crassipes Tchesunov & Mokievsky, 1995
Species: Batillipes dicrocercus Pollock, 1970
Species: Batillipes friaufi Riggin, 1962
Species: Batillipes gilmartini McGinty, 1969
Species: Batillipes lesteri Kristensen & Mackness, 2000
Species: Batillipes littoralis Renaud-Debyser, 1959
Subspecies: Batillipes littoralis littoralis Renaud-Debyser, 1959
Subspecies: Batillipes littoralis submersus d'Hondt, 1970
Species: Batillipes longispinosus Chang & Rho, 1997
Species: Batillipes marcelli Morone De Lucia, D'Addabbo Gallo and De Zio Grimaldi, 1988
Species: Batillipes mirus Richters, 1909
Synonym: Batillipes caudatus Hay, 1917
Species: Batillipes noerrevangi Kristensen, 1978
Species: Batillipes orientalis Chang & Rho, 1997
Species: Batillipes pennaki Marcus, 1946
Species: Batillipes philippinensis Chang & Rho, 1997
Species: Batillipes phreaticus Renaud Debyser, 1959
Synonym: Batillipes littoralis submersus D'Hondt, 1970
Species: Batillipes roscoffensis Kristensen, 1978
Species: Batillipes rotundiculus Rho, Min & Chang, 1999
Species: Batillipes similis Schulz, 1955
Species: Batillipes spinicauda Gallo D’Addabbo, Sandulli & de Zio Grimaldi, 2005
Species: Batillipes tridentatus Pollock, 1989
Species: Batillipes tubernatis Pollock, 1971
Familia: Coronarctidae Renaud-Mornant, 1974
Genus: Coronarctus Renaud-Mornant, 1974
Species: Coronarctus disparilis Renaud-Mornant, 1987
Species: Coronarctus fastigatus Renaud-Mornant, 1987
Species: Coronarctus laubieri Renaud-Mornant, 1987
Species: Coronarctus mexicus Romano III, Gallo, D'Addabbo, Accogli, Baguley & Montagna, 2011
Species: Coronarctus stylisetus Renaud-Mornant, 1987
Species: Coronarctus tenellus Renaud-Mornant, 1974
Species: Coronarctus verrucatus Hansen, 2007
Genus: Trogloarctus Villora-Moreno, 1996
Species: Trogloarctus trionyches Villora-Moreno, 1996
Familia: Halechiniscidae Thulin, 1928
Subfamilia: Dipodarctinae Pollock, 1995
Genus: Dipodarctus Pollock, 1995
Species: Dipodarctus anaholiensis Pollock, 1995
Species: Dipodarctus borrori Pollock, 1995
Synonym: Hemitanarctus chimaera de Zio Grimaldi, D'Addabbo Gallo,Morone De Lucia & Troccoli, 1995/96
Species: Dipodarctus subterraneus Renaud-Debyser, 1959


Subfamilia: Euclavarctinae Renaud-Mornant, 1983
Genus: Clavarctus Renaud-Mornant, 1983
Species: Clavarctus falculus Renaud-Mornant, 1983
Genus: Euclavarctus Renaud-Mornant, 1975
Species: Euclavarctus convergens Renaud-Mornant, 1983
Species: Euclavarctus thieli Renaud-Mornant, 1975
Genus: Exoclavarctus Renaud-Mornant, 1983
Species: Exoclavarctus dineti Renaud-Mornant, 1983
Genus: Moebjergarctus Bussau, 1992
Species: Moebjergarctus manganis Bussau, 1992
Genus: Parmursa Renaud-Mornant, 1984
Species: Parmursa fimbriata Renaud-Mornant, 1984
Species: Parmursa torquata Hansen, 2007
Genus: Proclavarctus Renaud-Mornant, 1983
Species: Proclavarctus fragilis Renaud-Mornant, 1983
Subfamilia: Florarctinae Renaud-Mornant, 1982
Genus: Florarctus Delamare Deboutteville & Renaud-Mornant, 1965
Species: Florarctus acer Renaud-Mornant, 1989
Species: Florarctus antillensis Van der Land, 1968
Species: Florarctus asper Renaud-Mornant, 1989
Species: Florarctus cervinus Renaud-Mornant, 1987
Species: Florarctus cinctus Renaud-Mornant, 1976
Species: Florarctus glareolus Noda, 1987
Species: Florarctus heimi Renaud-Mornant, 1965
Species: Florarctus hulingsi Renaud-Mornant, 1976
Species: Florarctus kwoni Chang & Rho, 1997
Species: Florarctus pulcher Grimaldi de Zio, Lamarca, D’addabbo Gallo & Pietanza, 1999
Species: Florarctus salvati Renaud-Mornant, 1965
Species: Florarctus stellatus Renaud-Mornant, 1989
Species: Florarctus vulcanius Renaud-Mornant, 1987
Genus: Ligiarctus Renaud-Mornant, 1982
Species: Ligiarctus eastwardi Renaud-Mornant, 1982
Genus: Wingstrandarctus Kristensen, 1984
Species: Wingstrandarctus corallinus Kristensen, 1984
Species: Wingstrandarctus crypticus Renaud-Mornant, 1989
Species: Wingstrandarctus intermedius Renaud-Mornant, 1967
Subfamilia: Halechiniscinae Thulin, 1928
Genus: Chrysoarctus Renaud-Mornant, 1984
Species: Chrysoarctus briandi Renaud-Mornant, 1984
Species: Chrysoarctus flabellatus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella & Grimaldi, 1982
Genus: Halechiniscus Richters, 1908


Species: Halechiniscus chafarinensis Grimaldi de Zio & Villora Moreno, 1995
Species: Halechiniscus greveni Renaud-Mornant & Deroux, 1976
Species: Halechiniscus guiteli Richters, 1908
Species: Halechiniscus jejuensis Chang & Rho, 2002
Species: Halechiniscus macrocephalus Grimaldi de Zio, D’Addabbo Gallo & Morone De Lucia, 1988
Species: Halechiniscus paratuleari Grimaldi de Zio, D’Addabbo Gallo & Morone De Lucia, 1988
Species: Halechiniscus perfectus Schulz, 1955
Species: Halechiniscus remanei Schulz, 1955
Subspecies: Halechiniscus remanei remanei Schulz, 1955
Subspecies: Halechiniscus remanei antillensis Renaud-Mornant, 1984
Species: Halechiniscus tuleari Renaud-Mornant, 1979
Genus: Paradoxipus Kristensen & Higgins, 1989
Species: Paradoxipus orzeliscoides Kristensen & Higgins, 1989
Subfamilia: Orzeliscinae Schulz, 1963
Genus: Orzeliscus du Bois-Reymond Marcus, 1952
Species: Orzeliscus belopus du Bois-Reymond Marcus, 1952
Species: Orzeliscus septentrionalis Schulz, 1953
Genus: Opydorscus Renaud-Mornant, 1989
Species: Opydorscus fonsecae Renaud-Mornant, 1989
Subfamilia: Styraconyxinae Kristensen & Renaud-Mornant, 1983
Genus: Angursa Pollock, 1979
Species: Angursa antarctica Villora-Moreno, 1998
Species: Angursa bicuspis Pollock, 1979
Subspecies: Angursa bicuspis bicuspis Pollock, 1979
Subspecies: Angursa bicuspis clavifera Noda, 1985
Subspecies: Angursa bicuspis abyssalis Renaud-Mornant, 1981
Species: Angursa capsula Bussau, 1992
Species: Angursa lanceolata Renaud-Mornant, 1981
Species: Angursa lingua Bussau, 1992
Genus: Bathyechiniscus Steiner, 1926
Species: Bathyechiniscus tetronyx Steiner, 1926
Genus: Lepoarctus Kristensen & Renaud-Mornant, 1983
Species: Lepoarctus coniferus Renaud-Mornant, 1975
Genus: Paratanarctus D'Addabbo Gallo, Grimaldi de Zio, Morone De Lucia & Troccoli, 1992
Species: Paratanarctus kristenseni D'Addabbo Gallo, Grimaldi de Zio, Morone De Lucia & Troccoli, 1992
Genus: Pleocola Cantacuzène, 1951
Species: Pleocola limnoriae Cantacuzène, 1951
Genus: Raiarctus Renaud-Mornant, 1981
Species: Raiarctus aureolatus Renaud-Mornant, 1981
Species: Raiarctus colurus Renaud-Mornant, 1981
Species: Raiarctus variabilis D’Addabbo Gallo, Grimaldi de Zio & Morone De Lucia, 1986
Genus: Rhomboarctus Renaud-Mornant, 1984
Species: Rhomboarctus aslaki Hansen, Gallo D’Addabbo & de Zio Grimaldi, 2003
Species: Rhomboarctus duplicicaudatus Hansen, Gallo D’Addabbo & de Zio Grimaldi, 2003
Species: Rhomboarctus thomassini Renaud-Mornant, 1984
Genus: Styraconyx Thulin, 1942
Species: Styraconyx craticuliformis Chang & Rho, 1998
Species: Styraconyx craticulus Pollock, 1983
Species: Styraconyx hallasi Kristensen, 1977
Species: Styraconyx haploceros Thulin, 1942
Species: Styraconyx kristenseni Renaud-Mornant, 1981
Subspecies: Styraconyx kristenseni kristenseni Renaud-Mornant, 1981
Subspecies: Styraconyx kristenseni neocaledonensis Renaud-Mornant, 1981
Species: Styraconyx nanoqsunguak Kristensen & Higgins, 1984
Species: Styraconyx paulae Robotti, 1971
Species: Styraconyx qivitoq Kristensen & Higgins, 1984
Species: Styraconyx sardiniae D'Addabbo Gallo, Morone De Lucia & Grimaldi de Zio, 1989
Species: Styraconyx sargassi Thulin, 1942
Species: Styraconyx testudo D'Addabbo Gallo, Grimaldi de Zio & Morone De Lucia, 1984
Species: Styraconyx tyrrhenus D'Addabbo Gallo, Morone De Lucia & Grimaldi de Zio, 1989
Genus: Tetrakentron Cuénot, 1892
Species: Tetrakentron synaptae Cuénot, 1892
Genus: Tholoarctus Kristensen & Renaud-Mornant, 1983
Species: Tholoarctus natans Kristensen & Renaud-Mornant, 1983
Subspecies: Tholoarctus natans natans Kristensen & Renaud-Mornant, 1983
Subspecies: Tholoarctus natans pedunculatus D'Addabbo Gallo, de Zio Grimaldi, Morone De Lucia & Troccoli, 1992
Subfamilia: Tanarctinae Renaud-Mornant, 1980
Genus: Actinarctus Schulz, 1935
Species: Actinarctus doryphorus Schulz, 1935
Subspecies: Actinarctus doryphorus doryphorus Schulz, 1935
Subspecies: Actinarctus doryphorus ocellatus Renaud-Mornant, 1971
Species: Actinarctus lyrophorus Renaud-Mornant, 1979
Species: Actinarctus neretinus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella & Grimaldi, 1982
Species: Actinarctus physophorus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella & Grimaldi, 1982
Genus: Tanarctus Renaud-Debyser, 1959
Species: Tanarctus arborspinosus Lindgren, 1971
Species: Tanarctus bubulubus Jørgensen & Kristensen, 2001
Species: Tanarctus dendriticus Renaud-Mornant, 1980
Species: Tanarctus gracilis Renaud-Mornant, 1980
Species: Tanarctus helleouetae Renaud-Mornant, 1984
Species: Tanarctus heterodactylus Renaud-Mornant, 1980
Species: Tanarctus longisetosus Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella & Grimaldi, 1982
Species: Tanarctus minotauricus Renaud-Mornant, 1984
Species: Tanarctus ramazzottii Renaud-Mornant, 1975
Species: Tanarctus tauricus Renaud-Debyser, 1959
Species: Tanarctus velatus McKirdy, Schmidt & McGinty-Bayly, 1976
Genus: Zioella Renaud-Mornant, 1987
Species: Zioella pavonina Renaud-Mornant, 1987
Familia: Neoarctidae De Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992
Genus: Neoarctus De Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992
Species: Neoarctus primigenius De Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992
Familia: Neostygarctidae De Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1982
Genus: Neostygarctus De Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1982
Species: Neostygarctus acanthophorus De Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1982
Familia: Renaudarctidae Kristensen & Higgins, 1984
Genus: Renaudarctus Kristensen & Higgins, 1984
Species: Renaudarctus psammocryptus Kristensen & Higgins, 1984
Familia: Stygarctidae Schulz, 1951
Subfamilia: Megastygarctidinae Bello & De Zio Grimaldi, 1999
Genus: Megastygarctides Schmidt & McGinty-Bayly, 1976
Species: Megastygarctides christinae Hansen & Kristensen, 2006
Species: Megastygarctides gerdae Hansen & Kristensen, 2006
Species: Megastygarctides isounguis Renaud-Mornant, 1981
Species: Megastygarctides orbiculatus Schmidt & McGinty-Bayly, 1976
Species: Megastygarctides setoloso Morgan & O'Reilly, 1988
Subfamilia: Stygarctinae Schulz, 1951
Genus: Parastygarctus Renaud-Debyser, 1965
Species: Parastygarctus biungulatus Morone De Lucia, Grimaldi de Zio & D'Addabbo Gallo, 1984
Species: Parastygarctus higginsi Renaud-Debyser, 1965
Species: Parastygarctus mediterranicus Gallo D’Addabbo, Grimaldi de Zio & Sandulli, 2001
Species: Parastygarctus renaudae Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia & Daddabbo, 1987
Species: Parastygarctus sterreri Renaud-Mornant, 1970
Genus: Pseudostygarctus McKirdy, Schmidt & McGinty-Bayly, 1976
Species: Pseudostygarctus apuliae Gallo D’Addabbo, de Zio Grimaldi & D’Addabbo, 2000
Species: Pseudostygarctus intermedius Renaud-Mornant, 1979
Species: Pseudostygarctus mirabilis de Zio Grimaldi, D’Addabbo Gallo & Morone De Lucia, 1998
Species: Pseudostygarctus rugosus Gallo D’Addabbo, de Zio Grimaldi & Sandulli, 2001
Species: Pseudostygarctus triungulatus McKirdy, Schmidt & McGinty-Bayly, 1976
Genus: Stygarctus Schulz, 1951
Species: Stygarctus abornatus McKirdy, Schmidt & McGinty-Bayly, 1976
Species: Stygarctus bradypus Schulz, 1951
Species: Stygarctus gourbaultae Renaud-Mornant, 1981
Species: Stygarctus granulatus Pollock, 1970
Species: Stygarctus lambertii Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia & Daddabbo, 1987
Species: Stygarctus spinifer Hiruta, 1985
Ordo: Echiniscoidea Richters, 1926
Familia: Carphanidae Binda and Kristensen, 1986
Genus: Carphania Binda, 1978
Species: Carphania fluviatilis Binda, 1978
Familia: Echiniscidae Thulin, 1928
Genus: Antechiniscus Kristensen, 1987
Species: Antechiniscus conversus Horning & Schuster, 1983
Species: Antechiniscus jermani Rossi & Claps, 1989
Species: Antechiniscus lateromamillatus Ramazzotti, 1964
Species: Antechiniscus moscali Claxton, 2001
Species: Antechiniscus parvisentus Horning & Schuster, 1983
Species: Antechiniscus perplexus Horning & Schuster, 1983
Genus: Bryochoerus Marcus, 1936
Species: Bryochoerus intermedius Murray, 1910
Subspecies: Bryochoerus intermedius intermedius Murray, 1910
Subspecies: Bryochoerus intermedius hawaiicus Thulin, 1928
Subspecies: Bryochoerus intermedius laevis Marcus, 1936
Genus: Bryodelphax Thulin, 1928
Species: Bryodelphax aaseae Kristensen, Michalczyk & Kaczmarek, 2010
Species: Bryodelphax alzirae du Bois-Reymond Marcus, 1944
Species: Bryodelphax amphoterus Durante Pasa & Maucci, 1975
Species: Bryodelphax asiaticus Kaczmarek & Michalczyk, 2004
Species: Bryodelphax atlantis Pilato & Lisi, 2008
Species: Bryodelphax brevidentatus Kaczmarek, Michalczyk & Degma, 2005
Species: Bryodelphax crossotus Grigarick, Schuster & Nelson, 1983
Species: Bryodelphax dominicanus Schuster & Toftner, 1982
Species: Bryodelphax iohannis Bertolani, Guidi & Rebecchi, 1996
Species: Bryodelphax lijiangensis Yang, 2002
Species: Bryodelphax mateusi Fontoura, 1982
Species: Bryodelphax meronensis Pilato, Lisi & Binda, 2010
Species: Bryodelphax ortholineatus Bartoš, 1963
Species: Bryodelphax parvulus Thulin, 1928
Species: Bryodelphax sinensis Pilato, 1974
Species: Bryodelphax tatrensis Weglarska, 1959
Species: Bryodelphax weglarskae Pilato, 1972
Genus: Cornechiniscus Maucci & Ramazzotti, 1981
Species: Cornechiniscus brachycornutus Maucci, 1987
Species: Cornechiniscus ceratophorus Maucci, 1973
Species: Cornechiniscus cornutus Richters, 1906
Synonym: Pseudechiniscus intermedius Mihelcic, 1970
Species: Cornechiniscus holmeni Petersen, 1951
Species: Cornechiniscus lobatus Ramazzotti, 1943
Synonym: Pseudechiniscus cornutus Mihelcic, 1966
Species: Cornechiniscus madagascariensis Maucci, 1993
Species: Cornechiniscus schrammi Dastych, 1979
Species: Cornechiniscus subcornutus Maucci & Ramazzotti, 1981
Species: Cornechiniscus tibetanus Maucci, 1979
Genus: Echiniscus C.A.S. Schultze, 1840
Species: Echiniscus africanus Murray, 1907
Species: Echiniscus aliquantillus Grigarick, Schuster & Nelson, 1983
Species: Echiniscus angolensis da Cunha & do Nascimento Ribeiro, 1964
Species: Echiniscus apuanus M. Bertolani, 1946
Species: Echiniscus arcangelii Maucci, 1973-74
Species: Echiniscus arctomys Ehrenberg, 1853
Species: Echiniscus arthuri Pilato, Binda & Lisi, 2005
Species: Echiniscus azoricus Pilato & Lisi, 2008
Species: Echiniscus baius Marcus, 1928
Species: Echiniscus baloghi Iharos, 1973
Species: Echiniscus barbarae Kaczmarek & Michalczyk, 2002
Species: Echiniscus batramiae Iharos, 1936
Species: Echiniscus becki Schuster & Grigarick, 1966
Species: Echiniscus bigranulatus Richters, 1907
Species: Echiniscus bisculptus Maucci, 1983
Species: Echiniscus blumi Richters, 1903
Synonym: Echiniscus ramazzottii Binda & Pilato, 1969
Subspecies: Echiniscus blumi blumi Richters, 1903
Subspecies: Echiniscus blumi schizofilus Bartoš, 1941
Species: Echiniscus calcaratus Richters, 1908
Species: Echiniscus calvus Marcus, 1931
Species: Echiniscus canadensis Murray, 1910
Synonym: Echiniscus punctulatus Mihelcic, 1955
Synonym: Echiniscus bellus Mihelcic, 1967
Species: Echiniscus canedoi da Cunha & do Nascimento Ribeiro, 1962
Species: Echiniscus capillatus Ramazzotti, 1956
Species: Echiniscus carsicus Mihelcic, 1966
Species: Echiniscus carusoi Pilato, 1972
Species: Echiniscus cavagnaroi Schuster & Grigarick, 1966
Species: Echiniscus cervicornis Murray, 1906
Species: Echiniscus charrua Claps & Rossi, 1997
Species: Echiniscus cheonyoungi Moon & Kim, 1994
Species: Echiniscus cirinoi Binda & Pilato, 1993
Species: Echiniscus clavispinosus Fontoura, Pilato & Lisi, 2011
Species: Echiniscus clevelandi Beasley, 1999
Species: Echiniscus columinis Murray, 1911
Species: Echiniscus corrugicaudatus McInnes, 2009
Species: Echiniscus crassispinosus Murray, 1907
Subspecies: Echiniscus crassispinosus crassispinosus Murray, 1907
Subspecies: Echiniscus crassispinosus fasciatus Marcus, 1928
Species: Echiniscus curiosus Claxton, 1996
Species: Echiniscus dariae Kaczmarek & Michalczyk, 2010
Species: Echiniscus darienae Miller, Horning & Dastych, 1995
Species: Echiniscus dearmatus Bartoš, 1935
Synonym: Echiniscus szaboi Iharos, 1973
Species: Echiniscus dikenli Maucci, 1973
Species: Echiniscus diploglyptus Durante Pasa & Maucci, 1975
Species: Echiniscus divergens Marcus, 1936
Species: Echiniscus dreyfusi de Barros, 1942
Species: Echiniscus duboisi Richters, 1902
Species: Echiniscus egnatiae Durante Pasa & Maucci, 1979
Species: Echiniscus ehrenbergi Kristensen, 1995
Species: Echiniscus elaeinae Pilato, Binda & Lisi, 2005
Species: Echiniscus elegans Richters, 1906
Species: Echiniscus evelinae de Barros, 1942
Species: Echiniscus filamentosus Plate, 1888
Species: Echiniscus ganczareki Michalczyk & Kaczmarek, 2007
Species: Echiniscus glaber Bartoš, 1937
Species: Echiniscus granulatus Doyère, 1840
Synonym: Echiniscus crassus Richters, 1904
Synonym: Echiniscus fortis Bartoš, 1935
Synonym: Echiniscus abanti Maucci, 1973
Species: Echiniscus heterospinosus Maucci, 1954
Species: Echiniscus hexacanthus Maucci, 1973
Species: Echiniscus hoonsooi Moon & Kim, 1990
Species: Echiniscus horningi Schuster & Grigarick, 1971
Species: Echiniscus inocelatus Mihelcic, 1938
Synonym: Echiniscus inocellatus Mihelcic, 1964
Species: Echiniscus insuetus Mihelcic, 1967
Species: Echiniscus jagodici Mihelcic, 1951
Species: Echiniscus jamesi Claxton, 1996
Species: Echiniscus japonicus Morikawa, 1951
Species: Echiniscus jenningsi Dastych, 1984
Species: Echiniscus kerguelensis Richters, 1904
Species: Echiniscus knowltoni Schuster & Grigarick, 1971
Species: Echiniscus kofordi Schuster & Grigarick, 1966
Species: Echiniscus kosickii Kaczmarek & Michalczyk, 2010
Species: Echiniscus lapponicus Thulin, 1911
Species: Echiniscus laterosetosus Ito, 1993
Species: Echiniscus laterospinosus Rudescu, 1964
Species: Echiniscus latifasciatus Dudichev & Biserov, 2000
Species: Echiniscus lichenorum Maucci, 1983
Species: Echiniscus limai da Cunha & do Nascimento Ribeiro, 1964
Species: Echiniscus lineatus Pilato, Fontoura, Lisi & Beasley, 2008
Species: Echiniscus longispinosus Murray, 1907
Species: Echiniscus loxophthalmus Richters, 1911
Species: Echiniscus madonnae Michalczyk & Kaczmarek, 2006
Species: Echiniscus maesi Séméria, 1985
Species: Echiniscus malpighii Biserov, 1994
Species: Echiniscus manuelae da Cunha & do Nascimento Ribeiro, 1962
Species: Echiniscus marcusi Pilato, Binda, 1989
Species: Echiniscus marginatus Binda & Pilato, 1994
Species: Echiniscus marginoporus Grigarick, Schuster & Nelson, 1983
Species: Echiniscus markezi Mihelcic, 1971/72
Species: Echiniscus marleyi X. Li, 2007
Species: Echiniscus mauccii Ramazzotti, 1956
Species: Echiniscus mediantus Marcus, 1930
Species: Echiniscus merokensis Richters, 1904
Synonym: Echiniscus iharosi Rudescu, 1964
Subspecies: Echiniscus merokensis merokensis Richters, 1904
Subspecies: Echiniscus merokensis suecicus Thulin, 1911
Species: Echiniscus migiurtinus Franceschi, 1957
Species: Echiniscus mihelcici Iharos, 1973
Species: Echiniscus militaris Murray, 1911
Species: Echiniscus molluscorum Fox & Garcia-Moll, 1962
Species: Echiniscus moniliatus Iharos, 1967
Species: Echiniscus montanus Iharos, 1982
Species: Echiniscus mosaicus Grigarick, Schuster & Nelson, 1983
Species: Echiniscus multispinosus da Cunha, 1944
Species: Echiniscus murrayi Iharos, 1969
Species: Echiniscus nelsonae X. Li, L. Wang & Yu, 2007
Species: Echiniscus nepalensis Dastych, 1975
Species: Echiniscus nigripustulus Horning, Schuster & Grigarick, 1978
Species: Echiniscus nobilis Mihelcic, 1967
Species: Echiniscus oihonnae Richters, 1903
Species: Echiniscus ollantaytamboensis Nickel, Miller & Marley, 2001
Species: Echiniscus osellai Maucci, 1975
Species: Echiniscus pajstunensis Bartoš,1941
Species: Echiniscus palmai Dastych, 1997
Species: Echiniscus perarmatus Murray, 1907
Species: Echiniscus peruvianus Binda & Pilato, 1994
Species: Echiniscus perviridis Ramazzotti, 1959
Species: Echiniscus phocae du Bois-Reymond Marcus, 1944
Species: Echiniscus polygonalis Ito, 1993
Species: Echiniscus pooensis Rodriguez-Roda, 1948
Species: Echiniscus porabrus Horning, Schuster & Grigarick, 1978
Species: Echiniscus postojnensis Mihelcic, 1967
Synonym: Echiniscus loxophtalmus Mihelcic, 1939
Species: Echiniscus pseudelegans Séméria, 1994
Species: Echiniscus pseudowendti Dastych, 1984
Species: Echiniscus punctus McInnes, 1995
Species: Echiniscus pusae Marcus, 1928
Species: Echiniscus quadrispinosus Richters, 1902
Subspecies: Echiniscus quadrispinosus quadrispinosus Richters, 1902
Synonym: Echiniscus scrofa Richters, 1902
Subspecies: Echiniscus quadrispinosus brachyspinosus Bartoš, 1934
Subspecies: Echiniscus quadrispinosus cribrosus Murray, 1907
Subspecies: Echiniscus quadrispinosus fissispinosus Murray, 1907
Species: Echiniscus quitensis Pilato, 2007
Species: Echiniscus rackae Dastych, 1986
Species: Echiniscus ranzii Ramazzotti, 1964
Species: Echiniscus reticulatus Murray, 1905
Species: Echiniscus reymondi Marcus, 1928
Species: Echiniscus robertsi Schuster & Grigarick, 1965
Species: Echiniscus rodnae Claxton, 1996
Species: Echiniscus rufoviridis du Bois-Reymond Marcus, 1944
Species: Echiniscus rugospinosus Marcus, 1928
Species: Echiniscus scabrospinosus Pilato et al. 2008b
Species: Echiniscus semifoveolatus Ito, 1993
Species: Echiniscus shaanxiensis X. Li, L. Wang & Yu, 2007
Species: Echiniscus siegristi Heinis, 1911
Species: Echiniscus simba Marcus, 1928
Species: Echiniscus speciosus Mihelcic, 1967
Synonym: Echiniscus roseus Mihelcic, 1967
Species: Echiniscus spiculifer Schaudinn, 1901
Species: Echiniscus spiniger Richters, 1904
Species: Echiniscus spinulosus Doyère, 1840
Species: Echiniscus storkani Bartoš, 1940
Species: Echiniscus sylvanus Murray, 1910
Species: Echiniscus taibaiensis L. Wang & X. Li, 2005
Species: Echiniscus tamus Mehlen, 1969 <Amended by Miller & Mehlen 2007>
Species: Echiniscus tardus Mihelcic, 1951
Species: Echiniscus tenuis Marcus, 1928
Species: Echiniscus tessellatus Murray, 1910
Species: Echiniscus testudo Doyère, 1840
Synonym: Echiniscus bellermanni C.A.S. Schultze, 1840
Synonym: Echiniscus inermis Richters, 1902
Synonym: Echiniscus trifilis Rahm, 1921
Synonym: Echiniscus filamentosus mongoliensis Iharos, 1973
Species: Echiniscus trisetosus Cuénot, 1932
Species: Echiniscus trojanus Maucci, 1973
Species: Echiniscus tropicalis Binda & Pilato, 1995
Species: Echiniscus tympanista Murray, 1911
Species: Echiniscus velaminis Murray, 1910
Species: Echiniscus vinculus Horning, Schuster & Grigarick, 1978
Species: Echiniscus virginicus Riggin, 1962 <Amended by Christenberry & Mason 1979>
Species: Echiniscus viridianus Pilato, Fontoura & Lisi, 2007
Species: Echiniscus viridis Murray, 1910 <Amended by Pilato et al. 2008a>
Species: Echiniscus viridissimus Péterfi, 1956
Species: Echiniscus walteri Pilato & Lisi, 2003
Species: Echiniscus weisseri Maucci, 1978
Species: Echiniscus wendti Richters, 1903
Species: Echiniscus zetotrymus Horning, Schuster & Grigarick, 1978
Genus: Hypechiniscus Thulin, 1928
Species: Hypechiniscus exarmatus Murray, 1907
Synonym: Oreella breviclava Grigarick, Schuster & Nelson, 1983
Species: Hypechiniscus gladiator Murray, 1905
Subspecies: Hypechiniscus gladiator gladiator Murray, 1905
Subspecies: Hypechiniscus gladiator bigladii Iharos, 1973
Subspecies: Hypechiniscus gladiator fissigladii Iharos, 1973
Subspecies: Hypechiniscus gladiator spinulosa Iharos, 1973
Species: Hypechiniscus papillifer Robotti, 1972
Genus: Mopsechiniscus du Bois-Reymond Marcus, 1944
Species: Mopsechiniscus frenoti Dastych, 1999
Species: Mopsechiniscus granulosus Mihelcic, 1967
Species: Mopsechiniscus imberbis Richters, 1907
Species: Mopsechiniscus schusteri Dastych, 1999
Species: Mopsechiniscus tasmanicus Dastych & Moscal, 1992
Genus: Novechiniscus Kristensen, 1987
Species: Novechiniscus armadilloides R.O. Schuster, 1975
Genus: Parechiniscus Cuénot, 1926
Species: Parechiniscus chitonides Cuénot, 1926
Species: Parechiniscus unispinosus da Cunha, 1947
Genus: Proechiniscus Kristensen, 1987
Species: Proechiniscus hanneae Petersen, 1951
Genus: Pseudechiniscus Thulin, 1911
Species: Pseudechiniscus alberti Dastych, 1987
Species: Pseudechiniscus asper Abe, Utsugi & Takeda, 1998
Species: Pseudechiniscus bartkei Weglarska, 1962
Species: Pseudechiniscus beasleyi X. Li, L. Wang & Yu, 2007
Species: Pseudechiniscus bidenticulatus Bartoš, 1963
Species: Pseudechiniscus bispinosus Murray, 1907
Species: Pseudechiniscus brevimontanus Kendall-Fite & Nelson, 1996
Species: Pseudechiniscus clavatus Mihelcic, 1955
Species: Pseudechiniscus conifer Richters, 1904
Species: Pseudechiniscus dicrani Mihelcic, 1938
Species: Pseudechiniscus distinctus Mihelcic, 1951
Species: Pseudechiniscus facettalis Petersen, 1951
Synonym: Pseudechiniscus pseudoconifer facettalis Maucci, 1954
Species: Pseudechiniscus goedeni Grigarick, Mihelcic & Schuster, 1964
Species: Pseudechiniscus gullii Pilato & Lisi, 2006
Species: Pseudechiniscus insolitus Maucci, 1991
Species: Pseudechiniscus islandicus Richters, 1904
Species: Pseudechiniscus jiroveci Bartoš, 1963
Species: Pseudechiniscus juanitae de Barros, 1939
Synonym: Pseudechiniscus suillus franciscae de Barros, 1942
Species: Pseudechiniscus jubatus Biserov, 1990
Species: Pseudechiniscus megacephalus Mihelcic, 1951
Species: Pseudechiniscus nataliae Biserov & Maucci, 1986
Species: Pseudechiniscus novaezeelandiae Richters, 1908
Subspecies: Pseudechiniscus novaezeelandiae novaezeelandiae Richters, 1908
Subspecies: Pseudechiniscus novaezeelandiae aspinosa Iharos, 1963
Subspecies: Pseudechiniscus novaezeelandiae laterospinosa Iharos, 1963
Subspecies: Pseudechiniscus novaezeelandiae marinae Bartoš, 1934
Species: Pseudechiniscus occultus Dastych, 1980
Species: Pseudechiniscus papillosus X. Li, L. Wang, Liu & Su, 2005
Species: Pseudechiniscus pilatoi X. Li, 2007
Species: Pseudechiniscus pseudoconifer Ramazzotti, 1943
Species: Pseudechiniscus pulcher Murray, 1910
Species: Pseudechiniscus quadrilobatus Iharos, 1969
Species: Pseudechiniscus ramazzottii Maucci, 1952
Subspecies: Pseudechiniscus ramazzottii ramazzottii Maucci, 1952
Subspecies: Pseudechiniscus ramazzottii facettalis Iharos, 1964
Synonym: Pseudechiniscus ramazzottii lineatus Maucci, 1973-74
Species: Pseudechiniscus raneyi Grigarick, Mihelcic & Schuster, 1964
Species: Pseudechiniscus santomensis Fontoura, Pilato & Lisi, 2010
Species: Pseudechiniscus scorteccii Franceschi, 1952
Species: Pseudechiniscus shilinensis Yang, 2002
Species: Pseudechiniscus sinensis Rahm, 1937
Species: Pseudechiniscus spinerectus Pilato, Binda, Napolitano & Moncada, 2001
Species: Pseudechiniscus suillus Ehrenberg, 1853
Synonym: Echiniscus mutabilis Murray, 1905
Species: Pseudechiniscus transsylvanicus Iharos, 1936
Species: Pseudechiniscus victor Ehrenberg, 1853
Synonym: Pseudechiniscus tridentifer Bartoš, 1935
Species: Pseudechiniscus yunnanensis L. Wang, 2009
Genus: Testechiniscus Kristensen, 1987
Species: Testechiniscus laterculus Schuster, Grigarick & Toftner, 1980
Species: Testechiniscus macronyx Richters, 1907
Species: Testechiniscus meridionalis Murray, 1906
Species: Testechiniscus spitsbergensis Scourfield, 1897


Synonym: Echiniscus clavisetosus Mihelcic, 1958
Synonym: Echiniscus marinellae Bartoš, 1935
Synonym: Echiniscus melanophthalmus Bartoš, 1936
Synonym: Echiniscus menzeli Heinis, 1917
Synonym: Echiniscus rosaliae Mihelcic, 1951
Synonym: Echiniscus spinuloides Murray, 1907
Familia: Echiniscoididae Kristensen & Hallas, 1980
Genus: Anisonyches Pollock, 1975
Species: Anisonyches deliquus Chang & Rho, 1998
Species: Anisonyches diakidius Pollock, 1975
Species: Anisonyches mauritianus De Zio Grimaldi, D'Addabbo Gallo, Morone De Lucia & D'Addabbo Gallo, 1987
Genus: Echiniscoides Plate, 1888
Species: Echiniscoides andamanensis Chang & Rho, 1998
Species: Echiniscoides bruni D'Addabbo Gallo, De Zio Grimaldi, Morone De Lucia & Troccoli, 1992
Species: Echiniscoides higginsi Hallas & Kristensen, 1982
Species: Echiniscoides hoepneri Kristensen & Hallas, 1980
Species: Echiniscoides horningi Kristensen, 1999
Species: Echiniscoides pollocki Hallas & Kristensen, 1982
Species: Echiniscoides sigismundi M. Schultze, 1865
Subspecies: Echiniscoides sigismundi galliensis Kristensen & Hallas, 1980
Subspecies: Echiniscoides sigismundi groenlandicus Kristensen & Hallas, 1980
Subspecies: Echiniscoides sigismundi hispaniensis Kristensen & Hallas, 1980
Subspecies: Echiniscoides sigismundi mediterranicus Kristensen & Hallas, 1980
Subspecies: Echiniscoides sigismundi polynesiensis Renaud-Mornant, 1976
Subspecies: Echiniscoides sigismundi porphyrae De Zio Grimaldi, D'Addabbo Gallo & Pietanza, 2000
Subspecies: Echiniscoides sigismundi sigismundi M. Schultze, 1865
Subspecies: Echiniscoides sigismundi verrucariae De Zio Grimaldi, D'Addabbo Gallo & Pietanza, 2000
Species: Echiniscoides travei Bellido & Bertrand, 1981
Familia: Oreellidae Puglia, 1959
Genus: Oreella Murray, 1910
Species: Oreella chugachii Calloway, Miller, Johansson & Whiting, 2011
Species: Oreella mollis Murray, 1910
Synonym: Oreella minor Ramazzotti, 1964
Species: Oreella vilucensis Rahm, 1931 (nomen dubium)
Synonym: Oreella bonnensis Rahm, 1931 (nomen dubium)
Classis: Mesotardigrada Rahm, 1937
Ordo: Thermozodia Ramazzotti and Maucci, 1983
Familia: Thermozodiidae Rahm, 1937
Genus: Thermozodium Rahm, 1937
Species: Thermozodium esakii Rahm, 1937