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- South China Sea Fisheries Research Institute, Chinese Academy of Fishery Science, Guangzhou 510300, China.
- Key Laboratory of Fishery Ecology and Environment of Guangdong Province, Guangzhou 510300, China.
- Du, F.Y., Xu, Z.Z., Huang, J.Q. & Guo, D.H. 2012. Studies on the medusae (Cnidaria) from the Beibu Gulf in the northern South China Sea, with description of three new species. Acta Zootaxonomica Sinica 37(3): 506–519. reference page