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- Ratcliffe, B.C., Smith, D.M. & Erwin, D. 2005. Oryctoantiquus borealis, new genus and species from the Eocene of Oregon, U.S.A., the world's oldest fossil dynastine and largest fossil scarabaeid (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists bulletin, 59: 127–135. PDF