Ralph W. Holzenthal

Ralph W. Holzenthal, U.S. entomologist.

• Department of Entomology, Fisheries, and Wildlife, Clemson University, Clemson, USA

Taxon names authored

(List may be incomplete)

• 75 taxon names authored by Ralph W. Holzenthal

Eponyms

(List may be incomplete)

• 1 eponym of Ralph W. Holzenthal

Publications

(List may be incomplete)

2002

• Holzenthal, R.W. & Cressa, C. 2002. The Trichoptera, caddisflies, of Venezuela: three new species and records of Atopsyche Banks (Hydrobiosidae). Studies on Neotropical Fauna and Environment 37: 133-143. DOI: 10.1076/snfe.37.2.133.8578. . Reference page. 

2004

• Holzenthal R.W., Andersen T. 2004: The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae). Zootaxa, 511: 1–80.

2006

• Holzenthal, R.W. & Robertson, D.R. 2006. Four new species of Contulma from South America (Trichoptera: Anomalopsychidae). Zootaxa 1355: 49–59. DOI: 10.11646/zootaxa.1355.1.3, PDF. Reference page. 
• Robertson, D.R. & Holzenthal, R.W. 2006. The Neotropical caddisfly genus Canoptila (Trichoptera: Glossosomatidae). Zootaxa 1272: 45–59. DOI: 10.11646/zootaxa.1272.1.2, PDF. Reference page. 

2007

• Holzenthal R.W., Blahnik R.J., Prather A.L., Kjer K.M. 2007: Order Trichoptera Kirby, 1813 (Insecta), Caddisflies. Zootaxa, 1668: 639–698.

2008

• Robertson, D.R. & Holzenthal, R.W. 2008. Two New Species and a New Record of Protoptila from Bolivia (Trichoptera: Glossosomatidae: Protoptilinae). Annals of the Entomological Society of America 101(3): 465–473. DOI: 10.1603/0013-8746(2008)101[465:TNSAAN]2.0.CO;2, ResearchGate Reference page. 

2010

• Johanson, K.A. & Holzenthal, R.W. 2010. The snail-case caddisfly subgenus Helicopsyche (Feropsyche) in Costa Rica, with the description of 3 new species (Trichoptera: Helicopsychidae). Zootaxa 2689: 37–47. Preview full article (PDF). Reference page. 
• Thomson, R.E. & Holzenthal, R.W. 2010. New Neotropical species of the genus Austrotinodes Schmid (Trichoptera: Ecnomidae). Zootaxa 2437: 38–50. Preview. Reference page. 

2011

• Hamilton, S.W. & Holzenthal, R.W. 2011. Twenty-four new species of Polycentropus (Trichoptera, Polycentropodidae) from Brazil. ZooKeys 76: 1–53. DOI: 10.3897/zookeys.76.790. . Reference page. 
• Holzenthal, R.W., Morse, J.C. & Kjer, K.M. 2011. Order Trichoptera Kirby, 1813. Pp 209–211 In Zhang, Z.-Q. (ed.) 2011. Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 1–237. Open access. Reference page. . (PDF) Reference page. 

2012

• Thomson, R.E. & Holzenthal, R.W. 2012. New species and records of Hydroptilidae (Trichoptera) from Venezuela. ZooKeys 185: 19–39. DOI: 10.3897/zookeys.185.2909. . Reference page. 

2015

• Thomson, R.E., Holzenthal, R.W. 2015: A revision of the Neotropical caddisfly genus Leucotrichia Mosely, 1934 (Hydroptilidae, Leucotrichiinae). Zookeys 499: 1-100. DOI: 10.3897/zookeys.499.8360. Reference page. 
• Muñoz-Quesada, F.J. & Holzenthal, R.W. 2015. Revision of the Neotropical species of the caddisfly genus Wormaldia McLachlan (Trichoptera: Philopotamidae). Zootaxa 3998(1): 1–138. DOI: 10.11646/zootaxa.3998.1.1. Preview (PDF) Full article (PDF) ISBN 978-1-77557-763-8 (paperback); ISBN 978-1-77557-764-5 (Online edition). Reference page. 

2016

• Holzenthal, R.W., Blahnik, R.J. & Calor, A.R. 2016. Three new species of Helicopsyche von Siebold (Trichoptera: Helicopsychidae) from Brazil: IN Living among dragons: A festschrift in honor to Dr. Angelo B. M. Machado on occasion of his 80th birthday. Zootaxa 4078(1): 344–353. DOI: 10.11646/zootaxa.4078.1.29. Full article (PDF) Reference page. 
• Holzenthal, R.W. & Rios-Touma, B. 2016. A new Ecuadorian species of the rare Neotropical caddisfly genus Amphoropsyche Holzenthal (Trichoptera, Leptoceridae). ZooKeys 640: 59–67. DOI: 10.3897/zookeys.640.10344. Reference page. 
• Rázuri-Gonzales, E. & Holzenthal, R.W. 2016. New synonyms in the highly diverse caddisfly genus Smicridea (Trichoptera, Hydropsychidae). ZooKeys 637: 21-31. DOI: 10.3897/zookeys.637.10148. Reference page. 

2017

• Holzenthal, R.W. & Calor, A.R. 2017. Catalog of the Neotropical Trichoptera (Caddisflies). ZooKeys 654: 1–566. DOI: 10.3897/zookeys.654.9516. Reference page. 
• Rázuri-Gonzales, E., Holzenthal, R.W. & Ríos-Touma, B. 2017. Two new species of the rare Neotropical caddisfly genus Amphoropsyche Holzenthal (Trichoptera, Leptoceridae). ZooKeys 707: 63—72. DOI: 10.3897/zookeys.707.20759. Reference page. 

2018

• Holzenthal., R.W., Blahnik, R.J. & Ríos-Touma, B. 2018. New species and a new genus of Philopotamidae (Trichoptera) from the Andes of Bolivia and Ecuador. ZooKeys 780: 89–108. DOI: 10.3897/zookeys.780.26977 Reference page. 
• Rázuri-Gonzales, E., Holzenthal, R.W. & Ríos-Touma, B. 2018. New Atanatolica species from Ecuador (Trichoptera, Leptoceridae). ZooKeys 793: 97–114. DOI: 10.3897/zookeys.793.26712 Reference page. 

2022

• Holzenthal, R.W., Blahnik, R.J. & Ríos-Touma, B. 2022. A new genus and new species of Ecuadorian Philopotamidae (Trichoptera). Zookeys 1117ː 95–122. DOI: 10.3897/zookeys.1117.86984 Reference page. 
• Houghton, D.C., DeWalt, R.E., Hubbard, T., Schmude, K.L., Dimick, J.J., Holzenthal, R.W., Blahnik, R.J. & Snitgen, J.L. 2022. Checklist of the caddisflies (Insecta, Trichoptera) of the Upper Midwest region of the United States. Pp 287–300 In Pauls, S.U., Thomson, R. & Rázuri-Gonzales, E. (eds.). Special Issue in Honor of Ralph W. Holzenthal for a Lifelong Contribution to Trichoptera Systematics. ZooKeys 1111ː 1–488. Reference page. . DOI: 10.3897/zookeys.1111.72345 Reference page. 
• Thomson, R.E., Frandsen, P.B. & Holzenthal, R.W. 2022. A preliminary molecular phylogeny of the family Hydroptilidae (Trichoptera): an exploration of combined targeted enrichment data and legacy sequence data. Pp 467–488 In Pauls, S.U., Thomson, R. & Rázuri-Gonzales, E. (eds.). Special Issue in Honor of Ralph W. Holzenthal for a Lifelong Contribution to Trichoptera Systematics. ZooKeys 1111ː 1–488. Reference page. . DOI: 10.3897/zookeys.1111.85361 Reference page. 

2025

• Rázuri-Gonzales, E. & Holzenthal, R.W. 2025. The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441–478. DOI: 10.3897/zookeys.1263.150396. . Reference page. 
• Thomson, R.E., Ríos-Touma, B. & Holzenthal, R.W. 2025. Additions to the genus Rhyacopsyche Müller, 1879 (Trichoptera, Hydroptilidae) in Ecuador. ZooKeys 1263: 531–543. DOI: 10.3897/zookeys.1263.148084. . Reference page. 

Biographies

• Pauls, S.U., Thomson, R. & Rázuri-Gonzales, E. (eds.). Special Issue in Honor of Ralph W. Holzenthal for a Lifelong Contribution to Trichoptera Systematics. ZooKeys 1111ː 1–488. Reference page.