Template:Willyard et al., 2021
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- Willyard, A., Gernandt, D.S., Cooper, B., Douglas, C., Finch, K., Karemera, H., Lindberg, E., Langer, S.K., Lefler, J., Marquardt, P. & Pouncey, D.L. 2021. Phylogenomics in the hard pines (Pinus subsection Ponderosae; Pinaceae) confirms paraphyly in Pinus ponderosa, and places Pinus jeffreyi with the California big cone pines. Systematic Botany 46: 538–561. DOI: 10.1600/036364421X16312067913435 PDF