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From Wikispecies
Lötters, S.
,
Debold, P.
,
Henle, K.
,
Glaw, F.
&
Kneller, M.
1997. Ein neuer Pfeilgiftfrosch aus der
Epipedobates pictus
-Gruppe vom Osthang der Cordillera Azul in Perú.
Herpetofauna. Weinstadt
19: 25–34.
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