Template:Boyle et al., 2015

• Boyle, J.H., Kaliszewska, Z.A., Espeland, M., Suderman, T.R., Fleming, J., Heath, A. & Pierce, N.E. 2014 (Online) 2015 (Print). Phylogeny of the Aphnaeinae: myrmecophilous African butterflies with carnivorous and herbivorous life histories. Systematic Entomology 40(1): 169–182. DOI: 10.1111/syen.12098 .  
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