Template:Borges, de Sousa & Manzanilla, 2006
- Borges, A., De Sousa, L. & Manzanilla, J. 2006. Description of a new Tityus species (Scorpiones: Buthidae) from Sierra de Portuguesa, western Venezuela, based on morphological and mitochondrial DNA evidence. Zootaxa 1107: 49–68. Abstract & excerpt