Template:Beaman, Anderson & Beaman, 2001
- Beaman, J.H., Anderson, C. & Beaman, R.S. 2001. The Plants of Mount Kinabalu, Volume 4: Dicotyledon Families Acanthaceae to Lythraceae. [Kota Kinabalu, Malaysia:] Natural History Publications (Borneo) in association with the Royal Botanic Gardens, Kew. ISBN 983-812-051-0