Template:Aymard, 2005

• Aymard C., G.A. 2005. Verbenaceae. Pp. 407–445 in Steyermark, J.A. (†), Berry, P.E., Yatskievych, K. & Holst, B.K. (eds.), Flora of the Venezuelan Guayana, Vol. 9, Rutaceae–Zygophyllaceae. Missouri Botanical Garden Press, St. Louis. ISBN 1-930723-47-4.  
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