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It has been suggested that Ponerorchis should be broadened to include Chusua, and that Hemipilia should include single `orphan' species of Ponerorchis and Habenaria.


  • Bateman, R.M., P.M. Hollingsworth, J. Preston, Y.B. Luo, A.M. Pridgeon and M.W. Chase 2003: Molecular phylogenetics and evolution of Orchidinae and selected Habenariinae (Orchidaceae). Botanical Journal of the Linnean Society 142 Issue 1, Pages 1–40 Abstract
  • Stewart, J. 1994: Ponerorchis. Alpine orchid gems from Japan. The Plantsman 1:29–35

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