Koeleria spicata subsp. spicata

Koeleria spicata subsp. spicata

Taxonavigation

Taxonavigation: Poales
Classification System: APG IV Superregnum: Eukaryota Cladus: Diaphoretickes Cladus: CAM Cladus: Archaeplastida Regnum: Plantae Cladus: Angiosperms Cladus: Monocots Cladus: Commelinids Ordo: Poales

Familia: Poaceae
Subfamilia: Pooideae
Tribus: Poeae
Subtribus: Aveninae
Genus: Koeleria
Species: Koeleria spicata
Subspecies: K. s. subsp. spicata

Name

Koeleria spicata (L.) Barberá, Quintanar, Soreng & P.M.Peterson subsp. spicata autonym

Synonyms

• Heterotypic
  • Avena flavescens var. virescens Regel in Bull. Soc. Imp. Naturalistes Moscou 41(II): 299 (1868 publ. 1869)
  • Avena mollis Michx. in Fl. Bor.-Amer. 1: 72 (1803), nom. illeg.
  • Avena squarrosa Schrank in Denkschr. Königl.-Baier. Bot. Ges. Regensburg 1(2): 7 (1818)
  • Avena tolucensis Kunth in F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 1: 148 (1816)
  • Avena virescens (Regel) Regel in Trudy Imp. S.-Peterburgsk. Bot. Sada 7: 635 (1880)
  • Briza toluccensis Kunth in F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 1: 148 (1816), in error.
  • Calamagrostis falklandiae Steud. in Syn. Pl. Glumac. 1: 192 (1854)
  • Deyeuxia subspicata Desv. in Mém. Soc. Agric. Angers 1: 164 (1831), pro syn.
  • Graphephorum shearii (Scribn.) Rydb. in Bull. Torrey Bot. Club 32: 602 (1905)
  • Koeleria alaskana (Nash) Chepinoga in Bot. Pacifica 12(2): 126 (2023)
  • Koeleria aristata Loisel. in Fl. Gall., ed. 2, 1: 66 (1828)
  • Koeleria canescens Torr. ex Trin. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math., Seconde Pt. Sci. Nat. 4(2, Bot.): 13 (1836), pro syn.
  • Koeleria oreophila subsp. colombiana (Finot) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 5 (2019)
  • Koeleria pseudomollis Chepinoga in Bot. Pacifica 12(2): 126 (2023)
  • Koeleria pseudomongolica Chepinoga in Bot. Pacifica 12(2): 126 (2023)
  • Koeleria spicata subsp. alaskana (Nash) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 6 (2019)
  • Koeleria spicata subsp. andina (Benth.) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 6 (2019)
  • Koeleria spicata subsp. dianthema (Louis-Marie) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. geghamensis (Gabrieljan) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. himalaica (Hultén) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. hultenii (Chrtek) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. kinabaluensis (Chrtek) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. mollis (Kunth) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. mongolica (Hultén) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. ovatipaniculata (Hultén) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 8 (2019)
  • Koeleria spicata subsp. virescens (Regel) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 9 (2019)
  • Koeleria spicata subsp. wrangelensis (V.V.Petrovsky) Barberá, Quintanar, Soreng & P.M.Peterson in Phytoneuron 2019-46: 9 (2019)
  • Koeleria virescens (Regel) Chepinoga in Bot. Pacifica 12(2): 126 (2023)
  • Koeleria wrangelensis (V.V.Petrovsky) Chepinoga in Bot. Pacifica 12(2): 126 (2023)
  • Melica triflora Bigelow in New England J. Med. Surg. 5: 334 (1816)
  • Rupestrina pubescens Prov. in Fl. Canad. 2: 689 (1862), nom. superfl.
  • Trisetaria spicata subsp. ovatipaniculata (Hultén) Banfi & Soldano in Atti Soc. Ital. Sci. Nat. Mus. Civico Storia Nat. Milano 135: 385 (1994 publ. 1996)
  • Trisetaria spicata subsp. pilosiglumis (Fernald) Banfi & Soldano in Atti Soc. Ital. Sci. Nat. Mus. Civico Storia Nat. Milano 135: 385 (1994 publ. 1996)
  • Trisetaria tolucensis (Kunth) Poir. in Encycl., Suppl. 5: 366 (1817)
  • Trisetum airoides f. depauperatum Neuman in Sver. Fl.: 755 (1901)
  • Trisetum alaskanum Nash in Bull. New York Bot. Gard. 2: 155 (1901)
  • Trisetum albidum Sodiro in Revista Colegio Nac. Vincento Rocafuerte 11(40-41): 86 (1930)
  • Trisetum americanum Gand. in Bull. Soc. Bot. France 49: 182 (1902)
  • Trisetum andinum Phil. in Linnaea 29: 93 (1858), nom. illeg.
  • Trisetum andinum Benth. in Pl. Hartw.: 261 (1846)
  • Trisetum argenteum Scribn. in Bull. Div. Agrostol. U.S.D.A. 11: 49 (1898), nom. illeg.
  • Trisetum biflorum Phil. in Anales Univ. Chile 43: 568 (1873), nom. illeg.
  • Trisetum brittonii Nash in Bull. New York Bot. Gard. 1: 437 (1900)
  • Trisetum congdonii Scribn. & Merr. in Bull. Torrey Bot. Club 29: 470 (1902)
  • Trisetum dianthemum (Louis-Marie) Finot in Contr. U.S. Natl. Herb. 48: 664 (2003)
  • Trisetum disjunctum Louis-Marie in Rhodora 30: 218 (1928)
  • Trisetum fedtschenkoi Henrard in Blumea 3: 425 (1940)
  • Trisetum formosanum Honda in Bot. Mag. (Tokyo) 41: 636 (1927)
  • Trisetum fuegianum Gand. in Bull. Soc. Bot. France 60: 28 (1913)
  • Trisetum geghamense Gabrieljan in Fl. Rastitel'nost' Rast. Res. Arm. 16: 11 (2007)
  • Trisetum gracile Sodiro in Revista Colegio Nac. Vincento Rocafuerte 11(40-41): 85 (1930), nom. illeg.
  • Trisetum groenlandicum Steud. in Syn. Pl. Glumac. 1: 228 (1854)
  • Trisetum interruptum E.Fourn. in Mexic. Pl. 2: 108 (1886), nom. illeg.
  • Trisetum kitadakense Honda in Bot. Mag. (Tokyo) 45: 297 (1931)
  • Trisetum labradoricum Steud. in Syn. Pl. Glumac. 1: 228 (1854)
  • Trisetum majus Rydb. in Bull. Colorado Agric. Exp. Sta. 100: 34 (1906)
  • Trisetum molle Kunth in Révis. Gramin. 1: 101 (1829)
  • Trisetum molle subsp. alaskanum (Nash) Rebr. in Arktichesk. Fl. SSSR 2: 100 (1964)
  • Trisetum mongolicum (Hultén) Peschkova, Tzvelev & Prob. in Zlaki Rossii: 168 (2019)
  • Trisetum mongolicum var. elatior (Krylov) Peschkova in Fl. Sibir. 2: 75 (1990)
  • Trisetum montanum var. pilosum Louis-Marie in Rhodora 30: 213 (1928)
  • Trisetum montanum var. shearii (Scribn.) Louis-Marie in Rhodora 30: 213 (1928)
  • Trisetum nivosum E.Fourn. in Mexic. Pl. 2: 107 (1886)
  • Trisetum ochrostachyum Phil. in Linnaea 33: 290 (1864)
  • Trisetum oreophilum subsp. colombianum Finot in Curr. Topics Pl. Biol. 11: 62 (2010)
  • Trisetum ovatipaniculatum (Hultén) Galushko, Tzvelev & Prob. in Zlaki Rossii: 168 (2019)
  • Trisetum pubiflorum Hack. in Oesterr. Bot. Z. 52: 187 (1902)
  • Trisetum seravschanicum Roshev. in Bot. Mater. Gerb. Glavn. Bot. Sada R.S.F.S.R. 3: 88 (1922)
  • Trisetum shearii Scribn. in Circ. Div. Agrostol. U.S.D.A. 30: 8 (1901)
  • Trisetum spicatum var. alaskanum (Nash) Louis-Marie in Rhodora 30: 239 (1928)
  • Trisetum spicatum subsp. alaskanum (Nash) Hultén in Svensk Bot. Tidskr. 53: 210 (1959)
  • Trisetum spicatum var. andinum (Benth.) Louis-Marie in Rhodora 30: 239 (1929)
  • Trisetum spicatum subsp. andinum (Benth.) Hultén in Svensk Bot. Tidskr. 53: 224 (1959)
  • Trisetum spicatum subsp. bolivianum Hultén in Svensk Bot. Tidskr. 53: 224 (1959)
  • Trisetum spicatum var. brittonii (Nash) Louis-Marie in Rhodora 30: 239 (1928)
  • Trisetum spicatum var. compactum (Lange) Louis-Marie in Probl. Biol. Veg. - La Trappe: 96 (1941)
  • Trisetum spicatum subsp. congdonii (Scribn. & Merr.) Hultén in Svensk Bot. Tidskr. 53: 218 (1959)
  • Trisetum spicatum var. congdonii (Scribn. & Merr.) Hitchc. in Proc. Biol. Soc. Washington 41: 160 (1928)
  • Trisetum spicatum var. dianthemum Louis-Marie in Rhodora 30: 239 (1928)
  • Trisetum spicatum subsp. dianthemum (Louis-Marie) Finot in Curr. Topics Pl. Biol. 11: 66 (2010)
  • Trisetum spicatum f. elatior Krylov in Fl. Altai Gov. Tomsk 3: 1612 (1914)
  • Trisetum spicatum f. flavicundum (Norm.) Dahl in A.Blytt, Haandb. Norges Fl.: 85 (1902)
  • Trisetum spicatum subsp. formosanum (Honda) Veldkamp in Gard. Bull. Singapore 36: 132 (1983)
  • Trisetum spicatum var. formosanum (Honda) Ohwi in J. Jap. Bot. 17: 443 (1941)
  • Trisetum spicatum var. fuegianum (Hack.) Louis-Marie in Rhodora 30: 239 (1928)
  • Trisetum spicatum subsp. himalaicum Hultén in Gard. Bull. Singapore 36: 135 (1983)
  • Trisetum spicatum var. himalaicum (Hultén) P.C.Kuo & Z.L.Wu in Fl. Reipubl. Popularis Sin. 9(3): 141 (1987), with incorrect basionym cited.
  • Trisetum spicatum subsp. hultenii Chrtek in Acta Univ. Carol., Biol. 1967: 101 (1968)
  • Trisetum spicatum f. interruptum (Norm.) Dahl in A.Blytt, Haandb. Norges Fl.: 85 (1902)
  • Trisetum spicatum subsp. kinabaluense Chrtek in Folia Geobot. Phytotax. 5: 447 (1970)
  • Trisetum spicatum var. kitadakense (Honda) Ohwi in J. Jap. Bot. 17: 443 (1941)
  • Trisetum spicatum var. laxius (Lange) Lindm. in O.R.Holmberg, Skand. Fl. 2: 175 (1926)
  • Trisetum spicatum var. luxius (Lange) Holmb. in Skand. Fl. 2: 175 (1926)
  • Trisetum spicatum var. maidenii (Gand.) Fernald in Rhodora 18: 196 (1916)
  • Trisetum spicatum var. majus (Rydb.) Farw. in Rep. (Annual) Michigan Acad. Sci. 21: 352 (1919 publ. 1920)
  • Trisetum spicatum subsp. majus (Rydb.) Hultén in Svensk Bot. Tidskr. 53: 218 (1959)
  • Trisetum spicatum var. michauxii H.St.John in Fl. S.-E. Washington: 62 (1937)
  • Trisetum spicatum var. molle (Kunth) Beal in Grass. N. Amer. 2: 377 (1896)
  • Trisetum spicatum subsp. molle (Kunth) Hultén in Svensk Bot. Tidskr. 53: 216 (1959)
  • Trisetum spicatum var. mongolicum (Hultén) P.C.Kuo & Z.L.Wu in Fl. Reipubl. Popularis Sin. 9(3): 140 (1987), with incorrect basionym cited.
  • Trisetum spicatum subsp. mongolicum Hultén in Gard. Bull. Singapore 36: 135 (1983)
  • Trisetum spicatum var. nivosum (E.Fourn.) Louis-Marie in Rhodora 30: 239 (1928)
  • Trisetum spicatum subsp. ovatipaniculatum Hultén in Svensk Bot. Tidskr. 69: 132 (1975)
  • Trisetum spicatum var. pilosiglume Fernald in Rhodora 18: 195 (1916)
  • Trisetum spicatum subsp. pilosiglume (Fernald) Hultén in Svensk Bot. Tidskr. 53: 215 (1959)
  • Trisetum spicatum var. pubiflorum (Hack.) L.Liu in Vasc. Pl. Hengduan Mount. 2: 2224 (1994)
  • Trisetum spicatum var. spicatiforme Hultén in Svensk Bot. Tidskr. 53: 220 (1959)
  • Trisetum spicatum subsp. tolucense (Kunth) Hultén in Svensk Bot. Tidskr. 53: 222 (1959)
  • Trisetum spicatum var. villosissimum (Lange) Louis-Marie in Rhodora 30: 239 (1928)
  • Trisetum spicatum subsp. virescens (Regel) Tzvelev in Novosti Sist. Vyssh. Rast. 7: 65 (1970 publ. 1971)
  • Trisetum spicatum f. viviparum A.E.Porsild in Publ. Bot. (Ottawa) 4: 9 (1974 publ. 1975)
  • Trisetum spicatum subsp. wrangelense V.V.Petrovsky in Novosti Sist. Vyssh. Rast. 15: 2 (1979)
  • Trisetum subspicatum var. breviglume Hack. in C.J.F.Skottsberg, Wiss. Ergebn. Schwed. Sudpol.-Exped. 4(4): 6 (1906)
  • Trisetum subspicatum var. compactum Lange in Consp. Fl. Groenland. 3: 164 (1880)
  • Trisetum subspicatum var. flavicundum Norm. in Christ. Vidensk.-Selsk. Forh. 1893 (16): 54 (1893)
  • Trisetum subspicatum var. fuegianum Hack. in C.J.F.Skottsberg, Wiss. Ergebn. Schwed. Sudpol.-Exped. 4(4): 6 (1906)
  • Trisetum subspicatum var. glabrifolium Macloskie in Rep. Princeton Univ. Exped. Patagonia, Botany 8(Suppl.): 49 (1915)
  • Trisetum subspicatum var. interruptum Norm. in Christ. Vidensk.-Selsk. Forh. 1893 (16): 54 (1893)
  • Trisetum subspicatum var. laxius Lange in Consp. Fl. Groenland. 3: 164 (1880)
  • Trisetum subspicatum f. maidenii Gand. in Bull. Soc. Bot. France 49: 182 (1902)
  • Trisetum subspicatum var. majus Vasey in Bull. Colorado Agric. Exp. Sta. 100: 34 (1906), pro syn.
  • Trisetum subspicatum var. molle (Kunth) A.Gray in Manual, ed. 2: 572 (1856)
  • Trisetum subspicatum var. villosissimum Lange in Consp. Fl. Groenland. 3: 164 (1880)
  • Trisetum tolucense (Kunth) Kunth in Révis. Gramin. 1: 101 (1829)
  • Trisetum triflorum (Bigelow) Á.Löve & D.Löve in Univ. Colorado Stud., Ser. Biol. 17: 7 (1965)
  • Trisetum triflorum subsp. molle (Kunth) Á.Löve & D.Löve in Univ. Colorado Stud., Ser. Biol. 17: 8 (1965)
  • Trisetum variabile É.Desv. in C.Gay, Fl. Chil. 6: 351 (1854)
  • Trisetum virescens (Regel) B.Fedtsch. in Izv. Imp. Bot. Sada Petra Velikago 14(Suppl. 2): 64 (1914), nom. illeg.
  • Trisetum wrangelense (V.V.Petrovsky) Prob. in Sosud. Rast. Sovet. Dal'nego Vostoka 1: 163 (1985)

Distribution

Native distribution areas:

Continental: Cosmopolitan Regional: Eurasia, Australia & America Afghanistan, Alaska, Alberta, Aleutian Is., Altay, Amur, Antipodean Is., Argentina Northeast, Argentina South, Arizona, Austria, Bolivia, Borneo, British Columbia, Buryatiya, California, Chatham Is., Chile Central, Chile North, Chile South, China North-Central, China South-Central, Chita, Colombia, Colorado, Connecticut, Dominican Republic, East European Russia, East Himalaya, Ecuador, Falkland Is., Finland, France, Germany, Greenland, Guatemala, Haiti, Iceland, Idaho, Inner Mongolia, Iran, Irkutsk, Italy, Japan, Kamchatka, Kazakhstan, Khabarovsk, Kirgizstan, Korea, Krasnoyarsk, Kuril Is., Labrador, Magadan, Maine, Manchuria, Manitoba, Massachusetts, Mexico Central, Mexico Gulf, Mexico Northeast, Mexico Southeast, Mexico Southwest, Michigan, Minnesota, Mongolia, Montana, Nepal, Nevada, New Brunswick, New Guinea, New Hampshire, New Mexico, New South Wales, New York, New Zealand South, Newfoundland, North Carolina, North Caucasus, North European Russia, Northwest Territories, Norway, Nova Scotia, Nunavut, Ontario, Oregon, Pakistan, Pennsylvania, Peru, Primorye, Qinghai, Québec, Rhode I., Sakhalin, Saskatchewan, South Dakota, Spain, Svalbard, Sweden, Switzerland, Tadzhikistan, Taiwan, Tennessee, Texas, Tibet, Transcaucasus, Tuva, Utah, Uzbekistan, Vermont, Victoria, Virginia, Washington, West Himalaya, West Siberia, Wisconsin, Wyoming, Xinjiang, Yakutiya, Yukon Introduced into: Great Britain, Kerguelen References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition

References

Primary references

• Barberá, P., Quintanar, A., Peterson, P.M., Soreng, R.J. & Romaschenko, K. 2019. New combinations, new names, typifications, and a new section, sect. Hispanica in Koeleria (Poeae, Poaceae). Phytoneuron 2019-46: 1–13. PDF Reference page.  46: 6

Additional references

• (2022). https://doi.org/10.11646/phytotaxa.532.1.7. epublication.

Links

• Govaerts, R. et al. 2025. Koeleria spicata subsp. spicata in Kew Science Plants of the World Online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2025 November 17. Reference page. 
• International Plant Names Index. 2025. Koeleria spicata subsp. spicata. Published online. Accessed: November 17 2025. Reference page.

Vernacular names

• English: Spike trisetum, Northern Oat-grass
• suomi: Tähkäkaura
• русский: Трищетинник колосистый