Jörn Köhler

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Jörn Köhler (born 1970), German herpetologist.

Taxon names authored

(List may be incomplete)


(List may be incomplete)


  • DIRKSEN, L., P.L. IBISCH, J. KÖHLER & W. BÖHME (1995): Zur Herpetofauna der semihumiden Samaipata-Region, Bolivien. II. Reptilien. Herpetofauna 17 (99): 15–28.
  • KÖHLER, J., P.L. IBISCH, L. DIRKSEN & W. BÖHME (1995): Zur Herpetofauna der semihumiden Samaipata-Region, Bolivien. I. Amphibien. Herpetofauna 17 (98): 13-24.
  • KÖHLER, J. (1995): Erstnachweis von Hyla marmorata, Leptodactylus syphax und Phrynopus peruvianus für Bolivien. Herpetofauna 17 (98): 6-8.
  • Köhler, J. & Jungfer, K.-H. 1995. Eine neue Art und ein Erstnachweis von Fröschen der Gattung Eleutherodactylus aus Bolivien. Salamandra 31(3): 149–156. PDF Reference page
  • KÖHLER, J., L. DIRKSEN, P.L. IBISCH, G. RAUER, D. RUDOLPH & W. BÖHME (1995): Zur Herpetofauna des Sehuencas-Bergregenwaldes im Carrasco-Nationalpark, Bolivien. Herpetofauna 17 (96): 12-25.


  • REICHLE, S. & J. KÖHLER (1996): Geographic distribution: Gymnophiona: Caecilia marcusi. Herpetological Review 27 (4): 208.
  • KÖHLER, J. & W. BÖHME (1996): Anuran amphibians from the region of Pre-Cambrian rock outcrops (inselbergs) in northeastern Bolivia, with a note on the gender of Scinax Wagler, 1830 (Hylidae). Revue française d'Aquariologie et Herpétologie 23 (3/4): 133-140.
  • REICHLE, S. & J. KÖHLER (1996): Erstnachweis von drei Froscharten für Bolivien: Hyla tritaeniata, Phyllomedusa palliata und Colostethus marchesianus. Herpetofauna 18 (101): 32-34.


  • REICHLE, S. & J. KÖHLER (1997): A new species of Eleutherodactylus (Anura: Leptodactylidae) from the Andean slopes of Bolivia. Amphibia-Reptilia 18 (4): 333-337.
  • KÖHLER, J. (1997): Geographic distribution: Anura: Lepidobatrachus laevis. Herpetological Review 28 (3): 156.
  • KÖHLER, J. (1997): Geographic distribution: Anura: Chacophrys pierottii. Herpetological Review 28 (3): 155-156.
  • KÖHLER, J., S. REICHLE & G. PETERS (1997): Advertisement calls of three species of Bufo (Amphibia: Anura: Bufonidae) from lowland Bolivia. Stuttgarter Beiträge zur Naturkunde Serie A 562: 1-8.
  • LÖTTERS, S., F. CASTRO, J. KÖHLER & R. RICHTER (1997): Notes on the distribution and color variation of poison frogs of the genus Phyllobates from western Colombia (Anura: Dendrobatidae). Revue française d'Aquariologie et Herpétologie 24 (1/2): 55-58.
  • KÖHLER, J., F. GLAW & M. VENCES (1997): Notes on the reproduction of Rhombophryne (Anura: Microhylidae) at Nosy Be, northern Madagascar. Revue française d'Aquariologie et Herpétologie 24 (1/2): 53-54.


  • KÖHLER, J., V. R. MORALES, S. LÖTTERS, S. REICHLE & J. APARICIO (1998): A new green species of frog, genus Eleutherodactylus, from Bolivia and Perú (Amphibia, Anura, Leptodactylidae). Studies on Neotropical Fauna and Environment 33 (2/3): 93-99.
  • KÖHLER, J. & S. REICHLE (1998): Geographic distribution: Anura: Cochranella pluvialis. Hepetological Review 29 (3): 172.
  • DE LA RIVA, I. & J. KÖHLER (1998): A new minute leptodactylid frog, genus Phyllonastes, from humid montane forests of Bolivia. Journal of Herpetology 32 (3): 325-229.
  • REICHLE, S. & J. KÖHLER (1998): Saisonale und wasserstandsabhängige Rufplatzverteilung von Froschlurchen der südlichen Beni-Savannen, Bolivien. Salamandra 34 (1): 43-54.
  • GLAW, F., J. KÖHLER, S. LÖTTERS & M. VENCES (1998): Preliminary list and references of newly described amphibian species and subspecies between 1993 and 1997. Elaphe 6 (1): 1-24.
  • KÖHLER, J., F. GLAW & M. VENCES (1998 "1997"): First record of Mabuya comorensis (Reptilia: Scincidae) for the Madagascan fauna, with notes on the reptile fauna of the offshore island Nosy Tanikely. Bollettino del Museo Regionale di Scienze Naturali Torino 15 (1): 75-82.
  • GLAW, F. & J. KÖHLER (1998): Amphibian species diversity exceeds that of mammals. Herpetological Review 29 (1): 11-12.


  • LÖTTERS, S., F. GLAW, S. REICHLE, J. KÖHLER & E. MEYER (1999): Notes on vocalizations in three species of Atelopus from Central and South America (Amphibia: Anura). Herpetozoa 12 (1/2): 79-83.
  • LÖTTERS, S., F. GLAW, J. KÖHLER & F. CASTRO (1999): On the geographic variation of the advertisement call of Dendrobates histrionicus Berthold, 1845 and related forms from northwestern South America (Anura: Dendrobatidae). Herpetozoa 12 (1/2): 23-38.
  • KÖHLER, J. & S. LÖTTERS (1999 "1998"): Annotated list of amphibian records from the Departamento Pando, Bolivia, with description of some advertisement calls. Bonner zoologische Beiträge 48 (3/4): 259-273.
  • KÖHLER, J. & S. LÖTTERS (1999): Advertisement calls of two Bolivian Leptodactylus (Amphibia: Anura: Leptodactylidae). Amphibia-Reptilia 20 (2): 215-219.
  • KÖHLER, J. & S. LÖTTERS (1999): New species of the Eleutherodactylus unistrigatus group (Amphibia: Anura: Leptodactylidae) from montane rain forest of Bolivia. Copeia 1999 (2): 422-427.
  • LÖTTERS, S., J. KÖHLER & S. REICHLE (1999): Description of the tadpole of the Andean treefrog Hyla marianitae (Amphibia: Anura: Hylidae). Folia Zoologica 48 (1): 49-53.
  • GLAW, F., M. VENCES, T. ZIEGLER, W. BÖHME & J. KÖHLER (1999): Specific distinctness and biogeography of the dwarf chameleons Brookesia minima, B. peyrierasi and B. tuberculata (Reptilia: Chamaeleonidae): evidence from hemipenial and external morphology. Journal of Zoology, London 247: 225-238.



  • KÖHLER, J. & S. LÖTTERS (2001): Description of a small tree frog, genus Hyla (Anura, Hylidae), from humid Andean slopes of Bolivia. Salamandra 37 (3): 175-184.
  • PADIAL, J. M. & J. KÖHLER (2001): First record of Physalaemus centralis (Anura: Leptodactylidae) for Bolivia, with description of its advertisement call. Boletín de la Asociación Herpetologica Española 12 (1): 6-8.
  • Ziegler, T. & Köhler, J. 2001. Rhacophorus orlovi sp. n., ein neuer Ruderfrosch aus Vietnam (Amphibia: Anura: Rhacophoridae). Sauria 23(3): 37–46. Reference page
  • KÖHLER, J. & S. LÖTTERS (2001): A new species of minute Hyla from the southwestern Amazon Basin (Amphibia, Anura, Hylidae). Studies on Neotropical Fauna and Environment 36 (2): 105-112.


  • ZIEGLER, T., H.-W. HERRMANN & J. KÖHLER (2002): Geographic distribution: Anura: Rhacophorus orlovi. Herpetological Review 33 (2): 146.


  • KÖHLER, J. (2003): Geographic distribution: Anura: Hyla cruzi. Herpetological Review 34 (4): 381.
  • KÖHLER, J., P. WAGNER, S. VISSER & W. BÖHME (2003): New country records of Adolfus africanus (Sauria: Lacertidae)  a rainforest lizard with disjunct distribution? Salamandra 39 (3/4): 241-248.


  • Böhme, W. & Köhler, J. 2005. Do endings of adjective flectible species names affect stability? A final note on the gender of Podarcis Wagler, 1830 (Reptilia: Lacertidae). Bonner zoologische Beiträge 47(3/4): 293–295. PDF Reference page
  • BÖHME, W., T. SCHÖTTLER, T. Q. NGUYEN & J. KÖHLER (2005): A new species of salamander, genus Tylototriton (Urodela: Salamandridae), from northern Vietnam. Salamandra 41 (4): 215-220.
  • Böhme, W., Wagner, P., Malonza, P., Lötters, S. & Köhler, J. 2005. A new species of the Agama agama group (Squamata: Agamidae) from western Kenya, East Africa, with comments on Agama lionotus Boulenger, 1896. Russian Journal of Herpetology 12(2): 143–150. DOI: 10.30906/1026-2296-2005-12-2-143-150 [nonfunctional] Broken access Reference page
  • KÖHLER, J., D. R. VIEITES, R. M. BONETT, F. HITA GARCIA, F. GLAW, D. STEINKE & M. VENCES (2005): New amphibians and global conservation: a boost in species discoveries in a highly endangered vertebrate group. BioScience 55 (8): 693-696. [incl. title page]
  • KÖHLER, J., K. SCHEELKE, S. SCHICK, M. VEITH & S. LÖTTERS (2005): Contribution to the taxonomy of hyperoliid frogs (Amphibia, Anura, Hyperoliidae): advertisement calls of twelve species from East and Central Africa. African Zoology 40 (1): 127-142.
  • DUELLMAN, W. E. & J. KÖHLER (2005): New species of marsupial frog (Hylidae: Hemiphractinae: Gastrotheca) from the Yungas of Bolivia. Journal of Herpetology 39 (1): 91-100.
  • Köhler, J., Jungfer, K.-H. & Reichle, S. 2005. Another new species of small Hyla (Anura, Hylidae) from Amazonian sub-Andean forest of western Bolivia. Journal of Herpetology 39(1): 43–50. DOI: 10.1670/0022-1511(2005)039[0043:ANSOSH]2.0.CO;2 Reference page


  • KÖHLER, J., B. BWONG, S. SCHICK, M. VEITH & S. LÖTTERS (2006): A new species of arboreal Leptopelis (Anura: Arthroleptidae) from the forests of western Kenya. Herpetological Journal 16 (2): 183-189.
  • MORAVEC, J., J. APARICIO & J. KÖHLER (2006): A new species of tree frog, genus Dendropsophus (Anura: Hylidae), from the Amazon of northern Bolivia. Zootaxa 1327: 23-40.
  • WOHLTMANN, A., J. KÖHLER & P. MARTIN (2006): Endoparasitic mite infections of anuran amphibians from Bolivian montane rain forests, with descriptions of two new Hannemania species (Acari: Parasitengona: Trombiculidae: Leeuwenhoekiinae). Organisms Diversity & Evolution 6 (2): 141-150.
  • PADIAL, J. M., J. KÖHLER & I. DE LA RIVA (2006): The taxonomic status of Hyla roeschmanni De Grys, 1938 (Anura: Hylidae). Zootaxa 1230: 63-68.
  • KÖHLER, J., A. JOHN & W. BÖHME (2006): Notes on amphibians recently collected in the Yungas de La Paz region, Bolivia. Salamandra 42 (1): 21-27.
  • FAIVOVICH, J., J. MORAVEC, D. F. CISNEROS-HEREDIA & J. KÖHLER (2006): A new species of the Hypsiboas benitezi group from the western Amazon basin (Amphibia: Anura: Hylidae). Herpetologica 62 (1): 96-108.


  • JANSEN, M. & J. KÖHLER (2007): The advertisement call of Eleutherodactylus fraudator (Anura: Brachycephalidae) from Bolivia. South American Journal of Herpetology 2 (3): 198-200.
  • KÖHLER, J., F. GLAW & M. VENCES (2007): A new green treefrog, genus Boophis Tschudi 1838 (Anura Mantellidae), from arid western Madagascar: phylogenetic relationships and biogeographic implications. Tropical Zoology 20 (2): 215-227.
  • KNOLL, A., J. KÖHLER, F. GLAW, M. TESCHKE & M. VENCES (2007): Larval morphology in four species of Madagascan frogs of the subgenus Brygoomantis (Mantellidae: Mantidactylus). Zootaxa 1616: 49-59.
  • PADIAL, J. M., S. CASTROVIEJO-FISHER, J. KÖHLER, E. DOMIC & I. DE LA RIVA (2007): Systematics of the Eleutherodactylus fraudator species group (Anura: Brachycephalidae). Herpetological Monographs 21: 213-240.
  • KÖHLER, J. & R. GÜSTEN (2007): Herpetological type specimens in the natural history collections of the museums in Darmstadt and Wiesbaden, Germany. Spixiana 30 (2): 275-288.
  • MORAVEC, J. & J. KÖHLER (2007): A new species of Chiasmocleis (Anura: Microhylidae) from the Iquitos region, Amazonian Peru, with possible direct development. Zootaxa 1605: 59-67.
  • WOHLTMANN, A., L. DU PREEZ, M.-O. RÖDEL, J. KÖHLER & M. VENCES (2007): Endoparasitic mites of the genus Endotrombicula Ewing, 1931 (Acari: Prostigmata: Parasitengona: Trombiculidae) from African and Madagascan anurans, with description of a new species. Folia Parasitologica 54: 225-235.
  • GLAW, F., J. KÖHLER, P. BORA, N.H.C RABIBISOA, O. RAMILIJAONA & M. VENCES (2007): Discovery of the genus Plethodontohyla (Anura: Microhylidae) in dry western Madagascar: description of a new species and biogeographic implications. Zootaxa 1577: 61-68.


  • ANDREONE, F., N. COX, F. GLAW, J. KÖHLER, N. A. C. RABIBISOA, H. RANDRIAMAHAZO, H. RANDRIANASOLO, C. J. RAXWORTHY, S. STUART, D. VALLAN & M. VENCES (2008): Update of the Global Amphibian Assessment for Madagascar in light of species discoveries, nomenclature changes, and new field information. Monografie del Museo Regionale di Scienze Naturali di Torino 45: 419-438.
  • D'CRUZE, N., J. KÖHLER, M. FRANZEN & F. GLAW (2008): A conservation assessment of the amphibians and reptiles of the Forêt d'Ambre Special Reserve, north Madagascar. Madagascar Conservation & Development 3 (1): 44-54.
  • MORAVEC, J., J. APARICIO, M. GUERRERO-REINHARD, G. CALDERON & J. KÖHLER (2008): Diversity of small Amazonian Dendropsophus (Anura: Hylidae): another new species from northern Bolivia. Zootaxa 1918: 1-12.
  • JANSEN, M. & J. KÖHLER (2008): Intraspecific combat behavior of Phyllomedusa boliviana (Anura: Hylidae) and the possible origin of visual signaling in nocturnal treefrogs. Herpetological Review 39 (3): 290-293.
  • WAGNER, P., J. KÖHLER, A. SCHMITZ & W. BÖHME (2008): The biogeographical assignment of a west Kenyan rain forest remnant: further evidence from analysis of its reptile fauna. Journal of Biogeography 35 (8): 1349-1361.
  • KÖHLER, J., F. GLAW & M. VENCES (2008): Two additional treefrogs of the Boophis ulftunni species group (Anura: Mantellidae) discovered in rainforests of northern and south-eastern Madagascar. Zootaxa 1814: 37-48.
  • ANDREONE, F., A. I. CARPENTER, N. COX, L. DU PREEZ, K. FREEMAN, S. FURRER, G. GARCIA, F. GLAW, J. GLOS, D. KNOX, J. KÖHLER, J. R. MENDELSON III, V. MERCURIO, R. A. MITTERMEIER, R. D. MOORE, N. H. C. RABIBISOA, H. RANDRIAMAHAZO, H. RANDRIANASOLO, N. RAMINOSOA, O. RAMILIJAONA, C. J. RAXWORTHY, D. VALLAN, M. VENCES, D. R. VIEITES & C. WELDON (2008): The challenge of conserving amphibian megadiversity in Madagascar. PLoS Biology 6 (5): e118. doi: 10.1371/journal.pbio.0060118.
  • RÖSLER, H., J. KÖHLER & W. BÖHME (2008): A new species of the diurnal gekkonid genus Pristurus Rüppell, 1835 from the Red Sea island Hanish al-Kabir, Yemen. Amphibia-Reptilia 29 (2): 217-227.
  • VENCES, M. & J. KÖHLER (2008): Global diversity of amphibians (Amphibia) in freshwater. Hydrobiologia 595: 569-580.
  • PADIAL, J. M., J. KÖHLER, A. MUÑOZ & I. DE LA RIVA (2008): Assessing the taxonomic status of tropical frogs through bioacoustics: geographical variation in the advertisement calls in the Eleutherodactylus discoidalis species group (Anura). Zoological Journal of the Linnean Society 152 (2): 353-365.


  • KÖHLER, J., D. R. VIEITES, F. GLAW, N. KAFFENBERGER & M. VENCES (2009): A further new species of limbless skink, genus Paracontias, from eastern Madagascar. African Journal of Herpetology 58 (2): 98-105.
  • Glaw, F., Köhler, J. & Vences, M. 2009. A distinctive new species of chameleon of the genus Furcifer (Squamata: Chamaeleonidae) from the Montagne d’Ambre rainforest of northern Madagascar. Zootaxa 2269: 32–42. Abstract & excerpt (PDF). Reference page
  • KNOLL, A., F. GLAW & J. KÖHLER (2009): The Malagasy snake Pseudoxyrhopus ambreensis preys upon chameleon eggs by shell slitting. Herpetology Notes 2: 161-162.
  • CROTTINI, A., J. DORDEL, J. KÖHLER, F. GLAW, A. SCHMITZ & M. VENCES (2009): A multilocus phylogeny of Malagasy scincid lizards elucidates the relationships of the fossorial genera Androngo and Cryptoscincus. Molecular Phylogenetics and Evolution 53: 345-350.
  • GLAW, F., J. KÖHLER & M. VENCES (2009): A new species of cryptically coloured day gecko (Phelsuma) from the Tsingy de Bemaraha National Park in western Madagascar. Zootaxa 2195: 61-68.
  • PADIAL, J. M., S. CASTROVIEJO-FISHER, J. KÖHLER, C. VILÀ, J. C. CHAPARRO & I. DE LA RIVA (2009): Deciphering the products of evolution at the species level: the need for an integrative taxonomy. Zoologica Scripta 38: 431-447.
  • PADIAL, J. M., J. C. CHAPARRO, J. KÖHLER & I. DE LA RIVA (2009): Rediscovery, resurrection and redescription of Rhinella leptoscelis (Boulenger, 1912) (Anura: Bufonidae). Zootaxa 2115: 56-64.
  • VIEITES, D. R., K. C. WOLLENBERG, F. ANDREONE, J. KÖHLER, F. GLAW & M. VENCES (2009): Vast underestimation of Madagascar's biodiversity evidenced by an integrative amphibian inventory. Proceedings of the National Academy of Science of the U.S.A. 106 (20): 8267-8272. [incl. title page]
  • GLAW, F., Z. T. NAGY, J. KÖHLER, M. FRANZEN & M. VENCES (2009): Phylogenetic relationships of a new species of pseudoxyrhophiine snake (Reptilia: Lamprophiidae: Thamnosophis) suggest a biogeographical link between western and northern Madagascar. Organisms Diversity & Evolution 9 (1): 13-22.
  • MEGSON, S., P. MITCHELL, J. KÖHLER, C. MARSH, M. FRANZEN, F. GLAW & N. D'CRUZE (2009): A comprehensive survey of amphibians and reptiles in the extreme north of Madagascar. Herpetology Notes 2: 31-44.
  • KÖHLER, J. (2009): The identity of Hylambates rufus aubryioides Andersson, 1907 (Anura: Arthroleptidae) from Cameroon. Copeia 2009 (1): 57-61.


  • VENCES, M., J. KÖHLER, A. CROTTINI & F. GLAW (2010): High mitochondrial sequence divergence meets morphological and bioacoustic conservatism: Boophis quasiboehmei sp. n., a new cryptic treefrog species from south-eastern Madagascar. Bonn zoological Bulletin 57 (2): 241–255.
  • KÖHLER, J., D. KOSCINSKI, J. M. PADIAL, J. C. CHAPARRO, P. HANDFORD, S. C. LOUGHEED & I. DE LA RIVA (2010): Systematics of Andean gladiator frogs of the Hypsiboas pulchellus species group (Anura, Hylidae). Zoologica Scripta 39 (6): 572-590.
  • VENCES, M., J. KÖHLER, M. PABIJAN & F. GLAW (2010): Two syntopic and microendemic new frogs of the genus Blommersia from the east coast of Madagascar. African Journal of Herpetology 59 (2): 133-156.
  • KÖHLER, J., M. VENCES, N. D'CRUZE & F. GLAW (2010): Giant dwarfs: discovery of a radiation of large-bodied 'stump-toed frogs' from karstic cave environments of northern Madagascar. Journal of Zoology 282 (1): 21-38.
  • GLAW, F., J. KÖHLER & M. VENCES (2010): A new fossorial frog, genus Rhombophryne, from Nosy Mangabe Special Reserve, Madagascar. Zoosystematics and Evolution 86 (2): 235-243.
  • GEHRING, P.-S., M. PABIJAN, F. M. RATSOAVINA, J. KÖHLER, M. VENCES & F. GLAW (2010): A Tarzan yell for conservation: a new chameleon, Calumma tarzan sp. n., proposed as a flagship species for the creation of new nature reserves in Madagascar. Salamandra 46 (3): 167–179.
  • D'CRUZE, N., J. KÖHLER, M. VENCES & F. GLAW (2010): A new fat fossorial frog (Microhylidae: Cophylinae: Rhombophryne) from rainforest of the Forêt d'Ambre Special Reserve, northern Madagascar. Herpetologica 66 (2): 182-191.
  • GLAW, F., P.-S. GEHRING, J. KÖHLER, M. FRANZEN & M. VENCES (2010): A new dwarf species of day gecko, genus Phelsuma, from the Ankarana pinnacle karst in northern Madagascar. Salamandra 46 (2): 83-92.
  • BORA, P., J. C. RANDRIANANTOANDRO, R. RANDRIANAVELONA, E. F. HANTALALAINA, R. R. ANDRIANTSIMANARILAFY, D. RAKOTONDRAVONY, O. R. RAMILIJAONA, M. VENCES, R. K. B. JENKINS, F. GLAW & J. KÖHLER (2010): Amphibians and reptiles of the Tsingy de Bemaraha plateau, western Madagascar: checklist, biogeography and conservation. Herpetological Conservation and Biology 5 (1): 111-125.
  • KÖHLER, J., M. VENCES, M. ERBACHER & F. GLAW (2010): Systematics of limbless scincid lizards from northern Madagascar: morphology, phylogenetic relationships and implications for classification (Squamata: Scincidae). Organisms Diversity & Evolution 10 (2): 147-159.
  • GLAW, F., J. KÖHLER, I. DE LA RIVA, D. R. VIEITES & M. VENCES (2010): Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis. Zootaxa 2383: 1-82.
  • SCHICK, S., B. ZIMKUS, A. CHANNING, J. KÖHLER & S. LÖTTERS (2010): Systematics of 'Little Brown Frogs' from East Africa: recognition of Phrynobatrachus scheffleri and description of a new species from the Kakamega Forest, Kenya (Amphibia: Phrynobatrachidae). Salamandra 46 (1): 24-36.
  • VENCES, M., F. GLAW, J. KÖHLER & K. C. WOLLENBERG (2010): Molecular phylogeny, morphology and bioacoustics reveal five additional species of arboreal microhylid frogs of the genus Anodonthyla from Madagascar. Contributions to Zoology 79 (1): 1-32.


  • KÖHLER, J., F. GLAW, G. M. ROSA, P.-S. GEHRING, M. PABIJAN, F. ANDREONE & M. VENCES (2011): Two new bright-eyed treefrogs of the genus Boophis from Madagascar. Salamandra 47 (4): 207–221.
  • WESENER, T., J. KÖHLER, S. FUCHS & D. VAN DEN SPIEGEL (2011): How to uncoil your partner - "mating songs" in giant pill-millipedes (Diplopoda: Sphaerotheriida). Naturwissenschaften 98 (11): 967–975.
  • PABIJAN, M. P.-S. GEHRING, J. KÖHLER, F. GLAW & M. VENCES (2011): A new microendemic frog species of the genus Blommersia from the east coast of Madagascar. Zootaxa 2978: 34–50.
  • GLAW, F., J. KÖHLER & M. VENCES (2011): New species of Gephyromantis from Marojejy National Park, northeast Madagasacar. Journal of Herpetology 45: 155–160.
  • VENCES, M., J. KÖHLER, D. R. VIEITES & F. GLAW (2011): Molecular and bioacoustic differentiation of deep conspecific lineages of the Malagasy treefrogs Boophis tampoka and B. luteus. Herpetology Notes 4: 239–246.
  • MIRALLES, A., J. KÖHLER, D. R. VIEITES, F. GLAW & M. VENCES (2011): Hypotheses on rostral shield evolution in fossorial lizards derived from the phylogenetic position of a new species of Paracontias (Squamata, Scincidae). Organisms Diversity & Evolution 11: 135-150.
  • RANDRIANIAINA, R. D., J. KÖHLER, J. GLOS, M. VENCES & F. GLAW (2011): Where to grow in the Tsingy? Limestone rock pools as breeding habitats of the relict frog Tsingymantis antitra from Madagascar and description of its tadpole. Salamandra 47 (2): 77–89.
  • Miralles, A., Köhler, J., Glaw, F. & Vences, M. 2011. A molecular phylogeny of the "Madascincus polleni species complex", with description of a new species of scincid lizard from the coastal dune area of northern Madagascar. Zootaxa 2876: 1–16. Preview Reference page


  • VENCES, M., M. GEHARA, J. KÖHLER & F. GLAW (2012): Description of a new Malagasy treefrog (Boophis) occurring syntopically with its sister species, and a plea for studies on non-allopatric speciation in tropical amphibians. Amphibia-Reptilia 33: 503–520.
  • RASMUSSEN, C., T. EISENBERG, D. ALFERMANN & J. KÖHLER (2012): Presence of Batrachochytrium dendrobatidis in amphibians from central and southern Hesse, central Germany: results from a preliminary regional screening. Salamandra 48 (3): 166–172.
  • ANDREONE, F., A. I. CARPENTER, J. COPSEY, A. CROTTINI, G. GARCIA, R. K. B. JENKINS, J. KÖHLER, N. H. C. RABIBISOA, H. RANDRIAMAHAZO & C. J. RAXWORTHY (2012): Saving the diverse Malagasy amphibian fauna: where are we four years after implementation of the Sahonagasy Action Plan? Alytes 29 (1-4): 44–58.
  • Glaw, F., Köhler, J. & Vences, M. 2012. A tiny new species of Platypelis from the Marojejy National Park in northeastern Madagascar (Amphibia: Microhylidae). European Journal of Taxonomy 9: 1–9. DOI: 10.5852/ejt.2012.9 Open access. Reference page
  • GLAW, F., J. KÖHLER, T. M. TOWNSEND & M. VENCES (2012): Rivaling the world's smallest reptiles: discovery of miniaturized and microendemic new species of leaf chameleons (Brookesia) from northern Madagascar. PLoS ONE 7(2): e31314. doi:10.1371/ journal.pone.0031314
  • KAFFENBERGER, N., K. C. WOLLENBERG, J. KÖHLER, F. GLAW, D. R. VIEITES & M. VENCES (2012): Phylogeny and biogeography of Malagasy frogs of the genus Gephyromantis. Molecular Phylogenetics and Evolution 62 (1): 555–560.






  • Rakotoarison, A., M. D. Scherz, F. Glaw, J. Köhler, F. Andreone, M. Franzen, J. Glos, O. Hawlitschek, T. Jono, A. Mori, S. H. Ndriantsoa, N. Rasoamampionona Raminosoa, J. C. Riemann, M.-O. Rödel, G. M. Rosa, D. R. Vieites, A. Crottini & M. Vences 2017. Describing the smaller majority: integrative taxonomy reveals twenty-six new species of tiny microhylid frogs (genus Stumpffia) from Madagascar. Vertebrate Zoology 67 (3): 271–398.
  • Scherz, M.D., Daza, J.D., Köhler, J., Vences, M. & Glaw, F. 2017. Off the scale: a new species of fish-scale gecko (Squamata: Gekkonidae: Geckolepis) with exceptionally large scales. PeerJ 5: e2955. DOI: 10.7717/peerj.2955 Open access Reference page





  • Castillo-Urbina, E., F. Glaw, C. Aguilar-Puntriano, M. Vences & J. Köhler (2021): Genetic and morphological evidence reveal another new toad of the Rhinella festae species group (Anura: Bufonidae) from the Cordillera Azul in central Peru. Salamandra 57 (2): 181–195.
  • Glaw, F., E. Castillo-Urbina, C. Aguilar-Puntriano, M. Vences & J. Köhler (2021): Anti-predator behaviour of the charismatic treefrog Cruziohyla craspedopus and its first record for central Peru (Amphibia, Phyllomedusidae). Spixiana 44 (2): 171–173.
  • Glaw, F., Köhler, J., Hawlitschek, O., Ratsoavina, F.M., Rakotoarison, A., Scherz, M.D. & Vences, M. 2021. Extreme miniaturization of a new amniote vertebrate and insights into the evolution of genital size in chameleons. Scientific Reports 11(1): 2522. DOI: 10.1038/s41598-020-80955-1 Open access Reference page
  • Glaw, F., J. Köhler, A. Crottini, P.-S. Gehring, D. Prötzel, L. Randriamanana, F. Andreone & M. Vences (2021): An additional level of cryptic diversity: a new green-coloured Malagasy treefrog of the Boophis luteus species group. Salamandra 57 (3): 295–308.
  • Miralles A., Bruy, T., Crottini, A., Rakotoarison, A., Ratsoavina, F.M., Scherz, M.D., Schmidt, R., Köhler, J., Glaw, F. & Vences, M. 2021. Completing a taxonomic puzzle: integrative review of geckos of the Paroedura bastardi species complex (Squamata, Gekkonidae). Vertebrate Zoology 71: 27–48. DOI: 10.3897/vz.71.e59495 Open access Reference page
  • Scherz, M. D., L. Schmidt, A. Crottini, A. Miralles, A. Rakotoarison, A. P. Raselimanana, J. Köhler, F. Glaw & M. Vences (2021): Into the Chamber of Horrors: A proposal for the resolution of nomenclatural chaos in the Scaphiophryne calcarata complex (Anura: Microhylidae), with a new species-level phylogenetic hypothesis for Scaphiophryninae. Zootaxa 4938 (1): 392–420.
  • Vences, M., J. Köhler, F. Andreone, A.-K. Craul, A. Crottini, L. du Preez, M. Preick, L. Rancilhac, M.-O. Rödel, M. D. Scherz, J. W. Streicher, M. Hofreiter & F. Glaw (2021): Target-enriched DNA sequencing clarifies the identity of name-bearing types of the Gephyromantis plicifer complex and reveals a new species of mantellid frog from Madagascar (Amphibia, Anura). Spixiana 44 (2): 175–202.


  • Köhler, J., Castillo-Urbina, E.F., Aguilar-Puntriano, C., Vences, M., Glaw, F. 2022. Rediscovery, redescription and identity of Pristimantis nebulosus (Henle, 1992), and description of a new terrestrial-breeding frog from montane rainforests of central Peru (Anura, Strabomantidae). Zoosystematics and Evolution 98(2): 213–232. DOI: 10.3897/zse.98.84963 Open access Reference page
  • Scherz, M. D., A. Crottini, C. R. Hutter, A. Hildenbrand, F. Andreone, T. R. Fulgence, G. Köhler, S. H. Ndriantsoa, A. Ohler, M. Preick, A. Rakotoarison, L. Rancilhac, A. P. Raselimanana, J. C. Riemann, M.-O. Rödel, G. M. Rosa, J. W. Streicher, D. R. Vieites, J. Köhler, M. Hofreiter, F. Glaw & M. Vences (2022): An inordinate fondness for inconspicuous brown frogs: integration of phylogenomics, archival DNA analysis, morphology, and bioacoustics yields 24 new taxa in the subgenus Brygoomantis (genus Mantidactylus) from Madagascar. Megataxa 7 (2): 113–311.
  • Vences, M., J. Köhler, A. Crottini, M. Hofreiter, C. R. Hutter, L. du Preez, M. Preick, A. Rakotoarison, L. Rancilhac, A. P. Raselimanana, G. M. Rosa, M. D. Scherz & F. Glaw (2022): An integrative taxonomic revision and redefinition of Gephyromantis (Laurentomantis) malagasius based on archival DNA analysis reveals four new mantellid frog species from Madagascar. Vertebrate Zoology 72: 271–309.



  • Gabriel, H., L.-D. Rothe, J. Köhler, S. Rakotomanga, D. Edmonds, P. Galán, F. Glaw, R. M. Lehtinen, A. Rakotoarison & M. Vences (2024): Unexpected diversity and co-occurrence of phytotelmic frogs (Guibemantis) around Andasibe, one of the most intensively surveyed amphibian hotspots of Madagascar, and description of three new species. Zootaxa 5397 (4): 451–485.
  • Köhler, J., F. Glaw, C. Aguilar-Puntriano, S. Castroviejo-Fisher, J. C. Chaparro, I. De la Riva, G. Gagliardi-Urrutia, R. Gutiérrez, M. Vences & J. M. Padial (2024): Similar looking sisters: A new sibling species in the Pristimantis danae group from the southwestern Amazon basin (Anura: Strabomantidae). Zoosystematics and Evolution 100 (2): 565–582.
  • Niessner, A. & J. Köhler (2024): Diurnal activity of an adult Ceratophrys cornuta preying on Copeoglossum nigropunctatum (Anura, Ceratophryidae and Squamata, Scincidae). Spixiana 46 (2): 320.
  • Vences, M., J. Köhler, M. D. Scherz, C. R. Hutter, H. M. Rabe Maheritafika, J. M. Rafanoharana, H. Raherinjatovo, A. Rakotoarison, F. Andreone, A. P. Raselimanana & F. Glaw (2024): Four new species of forest-dwelling mantellid frogs from Madagascar allied to Gephyromantis moseri (Amphibia, Anura). Spixiana 46 (2): 297–319.

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