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Taxonavigation: Scarabaeoidea 

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Scarabaeinae
Tribus: Onthophagini
Genus: Cyobius
Species (5): C. brendelli – C. cheyi – C. mendeli – C. tsukamotoi – C. wallacei


Cyobius Sharp in Harold, 1875


Additional references[edit]

  • Ochi, T.; Kon, M.; Barclay, M.V.L. 2010: Two new species of the genus Cyobius from Borneo (Coleoptera: Scarabaeidae). Entomological review of Japan, 65(2): 285–292. [Publication date: 'December 30, 2010', sourced from first page of article]
  • Ochi, T.; Kon, M.; Kashizaki, A. 2006: A new species of the genus Cyobius (Coleoptera, Scarabaeidae, Onthophagini) from Borneo. Elytra, Tokyo, 34(1): 145–152. [not seen]
  • Ochi, T.; Kon, M.; Kawahara, M. 2008: Notes on the coprophagous scarabaeid beetles (Coleoptera, Scarabaeidae) from Southeast Asia (XVII). A new species of Cyobius from Laos. Entomological review of Japan, 63(2): 175–179. [not seen]
  • Sharp 1875 in Harold, Coleopt. Hefte, 13, 48.


  • Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page.  [accessed on March 27, 2017]