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Taxonavigation: Adephaga 

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Adephaga

Familia: Dytiscidae
Subfamilia: Copelatinae
Genera (7): Agaporomorphus – Aglymbus – Copelatus – Exocelina – Lacconectus – Liopterus – Madaglymbus
[source: Michat & Torres (2009: 310)]


Copelatinae Van den Branden, 1885

Type genus[edit]


  • Balke, M.; Ribera, I.; Vogler, A.P. 2004: MtDNA phylogeny and biogeography of Copelatinae, a highly diverse group of tropical diving beetles (Dytiscidae). Molecular phylogenetics and evolution, 32: 866–880. DOI: 10.1016/j.ympev.2004.03.014
  • Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page
  • Michat, M.C.; Torres, P.L.M. 2009: A preliminary study on the phylogenetic relationships of Copelatus Erichson (Coleoptera: Dytiscidae: Copelatinae) based on larval chaetotaxy and morphology. Hydrobiologia, 632: 309–327. DOI: 10.1007/s10750-009-9853-2
  • Nilsson, A.N. 2007: Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998. Latissimus, 23: 33–34. ISSN: 0966-2235
  • Shaverdo, H.; Monaghan, M.T.; Lees, D.C.; Ranaivosolo, R.; Balke, M. 2008: Madaglymbus, a new genus of Malagasy endemic diving beetles and description of a highly unusual species based on morphology and DNA sequence data (Dytiscidae: Copelatinae). Systematics and biodiversity, 6: 43–51. DOI: 10.1017/S1477200007002599
  • Wewalka, G.; Balke, M.; Hendrich, L. 2010: Dytiscidae: Copelatinae (Coleoptera). Pp. 45-128 in: Jäch, M.A. & Balke, M. (eds.): Water beetles of New Caledonia (part 1). Monographs on Coleoptera, 3. ISSN: 1027-8869